Alignment with TRY

TRY is the plant trait database run by the Max Planck Institute for Biogeochemistry. It defines traits by name and number, such as trait 14, Leaf nitrogen (N) content per leaf dry mass. This page maps the 2661 TRY traits to BERVO. The work is tracked in issue #27 and is done in slices. The first slice was the map itself. The second added terms for the traits used by the plant-functional-traits schema. The third added the leaf economics and photosynthesis traits: leaf phosphorus, N:P ratios, photosynthetic and dark respiration rates, Vcmax, Jmax, stomatal conductance, leaf lifespan, thickness, tissue density, dry mass and phenology. The fourth added the organ and status concepts the broader rows named but BERVO lacked, such as twig, bark, coarse woody debris and senescent vegetation. The fifth added plant hydraulics and water relations: xylem vulnerability to embolism, leaf water potentials, hydraulic conductance and conductivity, the Huber value and xylem conduit size. The sixth added root traits: dry matter content, lignin content, respiration rate, branching ratio, root tip abundance, lifespan, rooting depth and mycorrhizal colonization, with the root to shoot ratio. The seventh added seed, fruit and dispersal traits: seed and diaspore dimensions, seed fresh mass, terminal velocity, oil and protein content, seeds per fruit, dispersal distance, fruit size and mass, and the base temperature and water potential for germination. The eighth added stem, wood and bark traits: bark thickness and density, with inner and outer bark, stem dry matter content, respiration rate and specific length, tree ring width, stem hydraulic conductivity, conduit lumen area and length, and sapwood area. The ninth added root traits that TRY also splits into coarse, fine, absorptive and transport roots: non-structural carbohydrate and starch content, stele diameter, cortex thickness, conduit diameter, specific root area and turnover rate, with organ dry mass per ground area mapped to the biomass terms. The tenth added potassium, calcium and magnesium content for any plant tissue, mapped the carbon, nitrogen and oxygen isotope signatures of plant organs to the existing isotope delta terms and the oxygen and hydrogen signatures of sapwood water to the water delta terms, and finished the woody organ biomass traits. The eleventh added the content of eleven more elements for any plant tissue (S, Na, Mn, Fe, Zn, Cu, Al, Ni, Si, B and Mo) and the most used leaf traits: stomatal density, leaf length and width, petiole length, fresh mass, chlorophyll, lignin, cellulose, non-structural carbohydrate and starch content, vein density, photosynthetic nitrogen and water use efficiency, transpiration rate and leaf area ratio. The twelfth added lignin, cellulose, hemicellulose, phenolics, non-structural carbohydrate and starch content for any plant tissue, with the leaf and root terms as their subclasses, and a decomposition rate constant for dead plant material. The thirteenth added, for any plant tissue, the lignin to nitrogen, carbon to phosphorus and potassium ratios, the soluble sugar, ash, protein, lipid, tannin and acid-insoluble residue contents, the lignin and acid-insoluble carbon fractions and the construction cost, and the fractions of mass, carbon, nitrogen and phosphorus remaining in decomposing plant material. The fourteenth added plant tissue pH, root length production and mortality, the necromass to biomass ratio, leaf isoprene and monoterpene emission rates, stomatal dimensions, the respiration rate and water content of any plant tissue and the nitrogen stock of vegetation, and mapped root mass production, mortality and necromass to the existing belowground production, belowground litter and vegetation biomass terms. The fifteenth added the vessel, fibre and parenchyma area fractions of wood, the dry matter content and modulus of elasticity of any plant tissue, relative bark thickness and the flexural, compressive and tensile strength of wood, and mapped twig and branch bark to the existing bark thickness and density terms, branch wood and bark C/N to the tissue C:N ratio, stem and sapwood water content to the tissue water content term, and petiole dry matter content to the tissue dry matter content term. The sixteenth added root length per ground area and per soil volume, root mass density, the carbon stock of vegetation, the root area index, root hair length and diameter, the stele diameter and area fractions of roots, root tips per unit root length and root acid phosphatase activity, with root hair and root tip concepts, and mapped root nitrogen per ground area to the vegetation nitrogen stock and fine root porosity to root porosity. The seventeenth added leaf carotenoid and anthocyanin contents, the thickness of the leaf epidermis, cuticle, mesophyll and its palisade and spongy layers, the light compensation and saturation points, leaf respiration in the light, nitrogen and phosphorus resorption efficiency, the maximum quantum efficiency of photosystem II, mesophyll conductance, the ratio of intercellular to ambient carbon dioxide and leaf equivalent water thickness. The eighteenth added, for any plant tissue, the bulk modulus of elasticity, the relative water content at and water lost to the turgor loss point, and the temperature sensitivity of respiration; leaf potassium and magnesium per unit area; seed germinability, viability, germination lag time, seed coat thickness, seed number per ground area and soil seed bank density; pollen grain diameter, pollen viability, and the pollen and seed to ovule ratios; flowers per plant and per inflorescence and fruits per infructescence; and the mass and nitrogen fractions of plant organs, the root length ratio, the age at first flowering and plant mortality rate, and mapped first flowering to the phenological event date. The nineteenth added the leaf maximum carboxylation and electron transport rates per unit nitrogen and their temperature sensitivities, the respiration rate per unit nitrogen of any plant tissue, the maximum PEP carboxylation rate, the triose phosphate utilisation rate, gross photosynthesis, the photosynthetic quantum yield, light response curvature and non-photochemical quenching; the mesophyll and chloroplast surface areas exposed to the intercellular space, leaf porosity, and leaf cell wall thickness, mass and content, with plant cell wall, chloroplast and intercellular space concepts; for any plant tissue, the water and osmotic potentials, the hydraulic capacitance and the osmotic solute content; the canopy radiation extinction coefficient; plant relative growth rate, lifespan, hydraulic conductance and nitrogen uptake efficiency; and the nitrogen and phosphorus content per seed and the numbers of seeds, fruits and inflorescences per plant and seeds per inflorescence. The trace element and contaminant contents wait on the decisions in issue #230. Still to come are the value lists of the categorical traits (woodiness, growth form, leaf type, leaf phenology) and terms for the gaps listed here.

Sources

Source	Revision	Used for
try_traits.csv, attached to issue #27	attached 2026-01-29, fetched 2026-10-22	The 2661 TRY traits, as trait_id and trait_name only. It is byte-identical to data/try_traits.csv in plant-functional-traits.
bioepic-data/plant-functional-traits, src/plant_functional_traits/schema/plant_functional_traits.yaml	d083f09	Which TRY traits the harmonized schema uses, in the pft_slot column

The TRY list has no units, definitions or value types. A term added from it takes those from the TRY data release documentation or from the literature the trait comes from, not from the list. TRY cuts trait names at 100 characters, so a few end mid-word; they are kept as given. Eleven names occur twice under different IDs, such as 456 and 1527, both Coarse root carbon (C) content per coarse root dry mass. The map keeps both IDs and points them at the same term.

The map

The map is src/scripts/try/try_alignment.tsv, written by src/scripts/build_try_alignment.py, one row per TRY trait:

Column	Holds
trait_id, trait_name	as in TRY
group	the organ or theme, used to plan slices
status	one of the five below
bervo_id	the BERVO term, for match, broader and related
pft_slot	the plant-functional-traits schema slot the trait feeds, if any
note	why a related term is not the same quantity, or another caveat

Status	Means
match	A BERVO term is the same quantity.
broader	A general BERVO term covers the trait, with what narrows it, such as an organ, a status or a time of day, in contexts. For example, Stem nitrogen (N) content per stem dry mass is Plant tissue nitrogen content with Stem in contexts.
related	A BERVO term is close but is not the same quantity, such as a reciprocal, a different normalisation, or an EcoSIM parameter defined differently. A term for the TRY trait is still needed.
gap	BERVO has no term.
species_attribute	The trait records where a species occurs, its habitat or indicator values, its legal status, its phylogeny or its uses, or it names a species, such as the dominant species of a plot or the herbivores of a leaf, rather than recording a property of a plant. These are set aside for a decision on scope, not rejected.

Groups and the species_attribute status are assigned by rules on the trait name, which were checked by hand. The C/N and N/P rows in broader are assigned the same way, as are the element contents per unit dry mass for C, N, P, K, Ca, Mg, S, Na, Mn, Fe, Zn, Cu, Al, Ni, Si, B and Mo, and the lignin, cellulose, hemicellulose, phenolics, non-structural carbohydrate and starch contents. Root contents per unit of soil mass and root microbial biomass were excluded from them, since neither is the content of a plant tissue. Every match and related row was assigned by hand.

A broader row is recorded with a concept for what narrows it in contexts: an organ, a status such as senescent, or a time of day such as predawn. Where the trait name says which, the row's note names the concept, such as Twig goes in contexts. Some TRY organs map to a BERVO concept with another name: a belowground plant organ is Belowground plant material, a reproductive organ is Reproductive structure, and senescent leaves take Senescent vegetation.

When an EcoSIM term is a match

Many BERVO plant terms are EcoSIM parameters with Plant functional type in contexts. Such a term is a match when it is the same quantity, as Seed length and Root porosity are. A value for a plant functional type is the trait's value for that type, as a TRY species mean is the trait's value for a species, and BERVO keeps one term per quantity whoever reports it. An EcoSIM variable resolved over something else, such as a branch, a node or a grid cell, measures a different thing and is related: Leaf number counts the leaves on a branch, not on a plant, and Leaf area is a grid-cell total.

How the alignment is recorded on a term

• A match term records the ID in its Comment as TRY trait ID: 14., or TRY trait IDs: 3108, 3109. when several TRY traits are the same quantity.
• A broader term records the IDs as TRY traits mapped here, with what narrows each one in contexts: 80, 406., following the pattern of the AmeriFlux BADM fields mapped to the same terms.
• A match term takes the TRY trait name as a Related Synonyms value, unless the name is already its label, is a label or synonym of another term, or is cut off at 100 characters in the TRY list. TRY's own abbreviations, such as SLA, LDMC and SSD, go in Exact Synonyms.
• TRY has no registered prefix, and its trait pages have no stable URL, so there is no TRY: cross-reference in DbXrefs.

just try-doc rebuilds this page, and it fails if the map and the Comment sentences disagree. tests/test_try_alignment.py runs it.

Summary

1146 TRY traits map to a BERVO term. 1463 are gaps and 52 are species attributes.

Group	match	broader	related	gap	species_attribute	Total
Crown and canopy	1	1	2	17		21
Fine root		166	24	213		403
Flower and reproduction	8	32	1	85		126
Leaf	70	55	36	256	1	418
Litter and woody debris	5	146	15	33		199
Moss and fern				21		21
Photosynthesis and gas exchange	40	7	17	49		113
Plot and stand	2			7	1	10
Root	31	135	17	225		408
Seed, fruit and dispersal	33	7	9	64		113
Species ecology and distribution				3	47	50
Species genetics and phylogeny				13	3	16
Stem, wood and bark	29	98	13	183		323
Twig, branch, shoot and bud		87	9	75		171
Whole plant	22	20	8	219		269
Total	241	754	151	1463	52	2661

Matches

TRY ID	TRY trait	BERVO term	Note
4	Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density	BERVO:0002371 Stem specific density
6	Root rooting depth	BERVO:0002418 Plant rooting depth
9	Root/shoot ratio	BERVO:0002414 Root to shoot ratio
12	Leaf lifespan (longevity)	BERVO:0002393 Leaf lifespan
13	Leaf carbon (C) content per leaf dry mass	BERVO:0002188 Leaf carbon content per unit dry mass
14	Leaf nitrogen (N) content per leaf dry mass	BERVO:0002189 Leaf nitrogen content per unit dry mass
15	Leaf phosphorus (P) content per leaf dry mass	BERVO:0002382 Leaf phosphorus content per unit dry mass
18	Plant height	BERVO:0002365 Plant height
21	Stem diameter	BERVO:0001884 Stem diameter
24	Bark thickness	BERVO:0002438 Bark thickness
26	Seed dry mass	BERVO:0002372 Seed dry mass
27	Seed length	BERVO:0000705 Seed length
37	Leaf phenology type	BERVO:0002397 Leaf phenology type
38	Plant woodiness	BERVO:0002377 Plant woodiness
40	Photosynthesis rate per leaf dry mass	BERVO:0002385 Leaf net photosynthetic rate per unit dry mass
41	Leaf respiration rate in the dark per leaf dry mass	BERVO:0002387 Leaf dark respiration rate per unit dry mass
42	Plant growth form	BERVO:0002378 Plant growth form	TRY variants differ in their value lists.
43	Leaf type	BERVO:0002379 Leaf type
45	Stomata conductance per leaf area	BERVO:0002392 Leaf stomatal conductance
46	Leaf thickness	BERVO:0002394 Leaf thickness
47	Leaf dry mass per leaf fresh mass (leaf dry matter content, LDMC)	BERVO:0002370 Leaf dry matter content
48	Leaf density (leaf tissue density, leaf dry mass per leaf volume)	BERVO:0002395 Leaf tissue density
50	Leaf nitrogen (N) content per leaf area	BERVO:0002187 Leaf nitrogen content per unit area
51	Leaf phosphorus (P) content per leaf area	BERVO:0002383 Leaf phosphorus content per unit area
52	Leaf potassium (K) content per leaf area	BERVO:0002567 Leaf potassium content per unit area
53	Photosynthesis rate per leaf area	BERVO:0002384 Leaf net photosynthetic rate per unit area
54	Leaf respiration rate in the dark per leaf area	BERVO:0002386 Leaf dark respiration rate per unit area
55	Leaf dry mass (single leaf)	BERVO:0002396 Individual leaf dry mass
59	Plant lifespan (longevity)	BERVO:0002613 Plant lifespan
63	Stomata density	BERVO:0002469 Stomatal density
66	Seed terminal velocity	BERVO:0002427 Diaspore terminal velocity	TRY names it for the seed; what falls is the diaspore.
77	Plant growth rate relative (plant relative growth rate, RGR)	BERVO:0002612 Plant relative growth rate
82	Root tissue density (root dry mass per root volume)	BERVO:0002376 Root tissue density
83	Root diameter	BERVO:0002375 Root diameter
87	Leaf lignin content per leaf dry mass	BERVO:0002476 Leaf lignin content per unit dry mass
88	Root lignin content per root dry mass	BERVO:0002412 Root lignin content per unit dry mass
92	Leaf cellulose content per leaf dry mass	BERVO:0002477 Leaf cellulose content per unit dry mass
95	Seed germination rate (germination efficiency)	BERVO:0002569 Seed germinability
96	Seed oil content per seed mass	BERVO:0002428 Seed oil content per unit mass
97	Seed protein content per seed mass	BERVO:0002429 Seed protein content per unit mass
101	Leaf epidermis thickness	BERVO:0002549 Leaf epidermis thickness
103	Leaf mesophyll thickness	BERVO:0002551 Leaf mesophyll thickness
109	Leaf area per plant dry mass (leaf area ratio; LAR)	BERVO:0002484 Leaf area ratio
111	Leaf transpiration rate per leaf area	BERVO:0002483 Leaf transpiration rate per unit area
112	Seed germination lag time	BERVO:0002571 Seed germination lag time
116	Root dry mass per root volume (root density, root tissue density)	BERVO:0002376 Root tissue density
123	Leaf intercellular volume per leaf volume	BERVO:0002602 Leaf porosity
131	Plant biomass and allometry: Seed number per plant	BERVO:0002618 Seed number per plant
134	Photosynthesis rate per leaf transpiration (photosynthetic water use effinciency: WUE)	BERVO:0002482 Photosynthetic water use efficiency
135	Photosynthesis rate per leaf nitrogen (N) content (photosynthetic nitrogen use efficiency, PNUE)	BERVO:0002481 Photosynthetic nitrogen use efficiency
143	Leaf petiole length	BERVO:0002472 Petiole length
144	Leaf length	BERVO:0002470 Leaf length
145	Leaf width	BERVO:0002471 Leaf width
155	Plant ontogeny: age of maturity (first flowering)	BERVO:0002585 Plant age at first flowering
163	Leaf fresh mass	BERVO:0002473 Individual leaf fresh mass
164	Leaf chlorophyll content per leaf dry mass	BERVO:0002475 Leaf chlorophyll content per unit dry mass
169	Stem conduit density (vessels and tracheids)	BERVO:0002410 Stem conduit density
170	Stem conduit cross-sectional area (vessels and tracheids)	BERVO:0002445 Stem conduit lumen area
171	Stem, branch or twig sapwood cross-sectional area per supported leaf area (Huber value)	BERVO:0002407 Huber value
183	Leaf epidermis cell wall thickness	BERVO:0002604 Leaf epidermis cell wall thickness
185	Photosynthesis carboxylation capacity (Vcmax) per leaf dry mass (Farquhar model)	BERVO:0002389 Leaf maximum carboxylation rate per unit dry mass
186	Photosynthesis carboxylation capacity (Vcmax) per leaf area (Farquhar model)	BERVO:0002388 Leaf maximum carboxylation rate per unit area
188	Leaf water & osmotic potential: leaf osmotic potential at full turgor	BERVO:0002401 Leaf osmotic potential at full turgor
189	Leaf water & osmotic potential: leaf osmotic potential at turgor loss	BERVO:0002400 Leaf water potential at turgor loss point	Osmotic and water potential are equal at turgor loss, where turgor is zero.
190	Leaf elastic modulus	BERVO:0002402 Leaf bulk modulus of elasticity
193	Dispersal distance	BERVO:0002431 Seed dispersal distance
210	Flower number of pollen per ovule	BERVO:0002577 Pollen to ovule ratio
233	Dispersal unit dry mass	BERVO:0002373 Diaspore dry mass
235	Dispersal unit thickness	BERVO:0002425 Diaspore thickness
236	Dispersal unit width	BERVO:0002424 Diaspore width
237	Dispersal unit length	BERVO:0002423 Diaspore length
238	Seed thickness	BERVO:0002422 Seed thickness
239	Seed width	BERVO:0002421 Seed width
246	Root porosity (fraction of root aerenchyma air space)	BERVO:0001077 Root porosity
248	Bark density (bark dry mass per bark volume)	BERVO:0002439 Bark density
269	Photosynthesis electron transport capacity (Jmax) per leaf area (Farquhar model)	BERVO:0002390 Leaf maximum electron transport rate per unit area
270	Photosynthesis electron transport capacity (Jmax) per leaf dry mass (Farquhar model)	BERVO:0002391 Leaf maximum electron transport rate per unit dry mass
281	Stem conduit diameter (vessels, tracheids)	BERVO:0002409 Stem conduit diameter
413	Leaf chlorophyll content per leaf area	BERVO:0002474 Leaf chlorophyll content per unit area
417	Leaf xanthophyll plus carotene content per leaf area	BERVO:0002545 Leaf carotenoid content per unit area
439	Plant respiration rate per plant nitrogen (N) content	BERVO:0002592 Plant tissue respiration rate per unit nitrogen	Whole plant; the term covers a whole plant.
486	Photosynthesis electron transport capacity (Jmax) per leaf nitrogen (N) content (Farquhar model)	BERVO:0002588 Leaf maximum electron transport rate per unit nitrogen
487	Photosynthesis carboxylation capacity (Vcmax) per leaf nitrogen (N) content (Farquhar model)	BERVO:0002587 Leaf maximum carboxylation rate per unit nitrogen
488	Photosynthesis electron transport capacity (Jmax) temperature dependence (Farquhar model)	BERVO:0002591 Leaf maximum electron transport rate temperature sensitivity
489	Crown (canopy) radiation extinction coefficient	BERVO:0002611 Canopy radiation extinction coefficient
491	Leaf carotenoid content per leaf area	BERVO:0002545 Leaf carotenoid content per unit area
505	Leaf starch content per leaf dry mass	BERVO:0002479 Leaf starch content per unit dry mass
507	Leaf non-structural carbon (TNC) content per leaf dry mass	BERVO:0002478 Leaf non-structural carbohydrate content per unit dry mass
510	Leaf nitrogen (N) retranslocation prior to leaf senescence	BERVO:0002557 Leaf nitrogen resorption efficiency
511	Plant number of branches per plant	BERVO:0000787 Number of branches of the plant
512	Leaf respiration rate in light per leaf area	BERVO:0002556 Leaf respiration rate in light per unit area
514	Root respiration rate per root dry mass	BERVO:0002413 Root respiration rate per unit dry mass
519	Stem respiration rate per stem dry mass	BERVO:0002441 Stem respiration rate per unit dry mass
533	Photosynthesis light response curvatur	BERVO:0002598 Leaf photosynthetic light response curvature
537	Photosynthesis carboxylation capacity (Vcmax) temperature dependence (Farquhar model)	BERVO:0002590 Leaf maximum carboxylation rate temperature sensitivity
570	Leaf carbon (C) content per leaf area	BERVO:0002186 Leaf carbon content per unit area
583	Leaf porosity	BERVO:0002602 Leaf porosity
668	Plant nitrogen (N) content per plant dry mass	BERVO:0002340 Plant tissue nitrogen content	Whole plant; the term covers whole plant material.
675	Root total non-structural carbon (TNC) content per root dry mass	BERVO:0002448 Root non-structural carbohydrate content per unit dry mass
709	Leaf cuticular conductance	BERVO:0002408 Leaf cuticular conductance
716	Leaf cuticula thickness	BERVO:0002550 Leaf cuticle thickness
719	Xylem hydraulic vulnerability, xylem cavitation vulnerability, embolism vulnerability, (P20, P50, P8	BERVO:0002398 Xylem vulnerability to embolism
720	Photosynthesis irradiance at saturation	BERVO:0002555 Leaf light saturation point
721	Photosynthesis light compensation point	BERVO:0002554 Leaf light compensation point
731	Leaf anthocyanin content per leaf dry mass	BERVO:0002548 Leaf anthocyanin content per unit dry mass
734	Stomata pore length	BERVO:0002518 Stomatal pore length
740	Leaf vein density	BERVO:0002480 Leaf vein density
748	Leaf hydraulic conductance	BERVO:0002404 Leaf hydraulic conductance
752	Plant hydraulic conductance	BERVO:0002614 Plant hydraulic conductance
770	Plant mortality rate	BERVO:0002586 Plant mortality rate
785	Root dry mass per root fresh mass (root dry matter content; RDMC)	BERVO:0002411 Root dry matter content
804	Leaf mesophyll palisade parenchym thickness	BERVO:0002552 Leaf palisade mesophyll thickness
805	Leaf mesophyll spongiophyll thickness	BERVO:0002553 Leaf spongy mesophyll thickness
809	Leaf carotenoid content per leaf dry mass	BERVO:0002546 Leaf carotenoid content per unit dry mass
810	Leaf anthocyanin content per leaf area	BERVO:0002547 Leaf anthocyanin content per unit area
917	Fruit width (diameter)	BERVO:0002433 Fruit width
918	Fruit length	BERVO:0002432 Fruit length
919	Fruit dry mass	BERVO:0002434 Fruit dry mass
928	Leaf area-specific conductivity (leaf specific conductivity)	BERVO:0002405 Leaf-specific hydraulic conductivity
938	Leaf isoprene emission rate per leaf area	BERVO:0002514 Leaf isoprene emission rate
964	Leaf mesophyll conductance	BERVO:0002560 Leaf mesophyll conductance
966	Leaf volatile organic carbon (VOC): monoterpene emission rate per leaf area	BERVO:0002515 Leaf monoterpene emission rate
982	Leaf relative water content (leaf water content fresh / leaf water content saturated; RWC)	BERVO:0002403 Leaf relative water content
983	Leaf water & osmotic potential: leaf water potential	BERVO:0002399 Leaf water potential
994	Litter decomposition rate constant	BERVO:0002491 Decomposition rate constant
1021	Plant carbon/nitrogen (C/N) ratio	BERVO:0002055 Plant tissue carbon to nitrogen mass ratio	Whole plant; the term covers a whole plant.
1031	Plant biomass and allometry: number of inflorescences per plant	BERVO:0002620 Inflorescence number per plant
1042	Pollen diameter	BERVO:0002575 Pollen grain diameter
1044	Pollen viability	BERVO:0002576 Pollen viability
1075	Root root hair length	BERVO:0002538 Root hair length
1080	Root length per root dry mass (specific root length, SRL)	BERVO:0002374 Specific root length
1085	Root persistence (lifespan, longevity)	BERVO:0002417 Root lifespan
1089	Root surface area per root dry mass (specific root area)	BERVO:0002453 Specific root area
1095	Wood (sapwood) cross-sectional area	BERVO:0002447 Sapwood area
1096	Wood (sapwood) specific conductivity (stem specific conductivity)	BERVO:0002406 Sapwood-specific hydraulic conductivity
1104	Inflorescence: seed number per inflorescence (total, fertile, infertile)	BERVO:0002621 Seed number per inflorescence
1105	Seed number per square meter	BERVO:0002572 Seed number per ground area
1109	Seed viability	BERVO:0002570 Seed viability
1111	Seedbank density	BERVO:0002573 Soil seed bank density
1177	Stem conduit length	BERVO:0002446 Stem conduit length
1181	Stem dry mass per stem fresh mass (stem dry matter content, StDMC)	BERVO:0002440 Stem dry matter content
1190	Stomata length	BERVO:0002516 Stomatal length
1192	Stomata width	BERVO:0002517 Stomatal width
1253	Leaf magnesium (Mg) content per leaf area	BERVO:0002568 Leaf magnesium content per unit area
1278	Root stele fraction: stele diameter divided by root diameter	BERVO:0002540 Root stele diameter fraction
1279	Root stele diameter	BERVO:0002450 Root stele diameter
1280	Root cross-sectional fraction of stele area to root area	BERVO:0002541 Root stele area fraction
1287	Root conduit diameter	BERVO:0002452 Root conduit diameter
1295	Root branching intensity 1 (branching ratio): number of roots per higher order root	BERVO:0002415 Root branching ratio
1298	Root tips per root dry mass (specific root tip abundance, SRTA)	BERVO:0002416 Specific root tip abundance
1401	Root growth: root length production per ground area per year	BERVO:0002511 Root length production
1418	Root dry mass turnover rate	BERVO:0002454 Root turnover rate
1430	Mycorrhizal colonization AM: fraction of root length that contains arbuscular mycorrhizal structures	BERVO:0002419 Arbuscular mycorrhizal root colonization
1431	Mycorrhizal colonization EM: fraction of root length colonized by ectomycorrhizal structure	BERVO:0002420 Ectomycorrhizal root colonization
1494	Root dry mass per soil volume	BERVO:0002535 Root mass density
1504	Plant biomass and allometry: Root length ratio (RLR): root length per plant dry mass	BERVO:0002584 Root length ratio
1507	Root length density (RLD): root length per ground area	BERVO:0002534 Root length per ground area
1508	Root length per soil volume	BERVO:0002544 Root length per soil volume
2700	Belowground plant organ growth: belowground plant organ dry mass production per ground area per year	BERVO:0002342 Belowground net primary production
2947	Inflorescence: number of flowers per inflorescence	BERVO:0002580 Flower number per inflorescence
2950	Paleo trait: leaf mass per area	BERVO:0002185 Leaf mass per area	Reconstructed from fossil leaves; the quantity is the same.
2999	Flower number per plant	BERVO:0002579 Flower number per plant
3002	Leaf mesophyll surface area exposed to internal airspace	BERVO:0002600 Leaf mesophyll surface area exposed to intercellular space per unit leaf area
3003	Leaf chloroplast surface area exposed to internal airspace	BERVO:0002601 Leaf chloroplast surface area exposed to intercellular space per unit leaf area
3018	Wood cross-sectional fraction of Vessel (conduit) area	BERVO:0002523 Wood vessel area fraction
3019	Wood cross-sectional fraction of fibre area	BERVO:0002524 Wood fibre area fraction
3020	Wood cross-sectional fraction of wod ray area	BERVO:0002525 Wood ray parenchyma area fraction
3021	Wood cross-sectional fraction of axial parenchyma area	BERVO:0002526 Wood axial parenchyma area fraction
3026	Leaf phosphorus (P) retranslocation prior to leaf senescence	BERVO:0002558 Leaf phosphorus resorption efficiency
3027	Plant carbon (C) content per plant dry mass	BERVO:0002339 Plant tissue carbon content	Whole plant; the term covers whole plant material.
3028	Plant phosphorus (P) content per plant dry mass	BERVO:0002380 Plant tissue phosphorus content	Whole plant; the term covers whole plant material.
3042	Seed volume	BERVO:0000704 Seed volume
3043	Seed surface area	BERVO:0000706 Seed surface area
3058	Leaf area (in case of compound leaves undefined if leaf or leaflet, petiole and rhachis excluded)	BERVO:0002368 Individual leaf area	TRY variants differ in whether the petiole is included.
3070	Plant nitrogen (N) uptake efficiency (plant nitrogen acquired per nitrogen available)	BERVO:0002615 Plant nitrogen uptake efficiency
3086	Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA) petiole, rhachis and midrib excluded	BERVO:0002369 Specific leaf area	TRY variants differ in whether the petiole, rhachis and midrib are included.
3106	Plant height vegetative	BERVO:0002366 Vegetative plant height
3107	Plant height generative	BERVO:0002367 Generative plant height
3108	Leaf area (in case of compound leaves: leaf, petiole excluded)	BERVO:0002368 Individual leaf area	TRY variants differ in whether the petiole is included.
3110	Leaf area (in case of compound leaves: leaf, petiole included)	BERVO:0002368 Individual leaf area	TRY variants differ in whether the petiole is included.
3112	Leaf area (in case of compound leaves: leaf, undefined if petiole in- or excluded)	BERVO:0002368 Individual leaf area	TRY variants differ in whether the petiole is included.
3114	Leaf area (in case of compound leaves undefined if leaf or leaflet, undefined if petiole is in- or e	BERVO:0002368 Individual leaf area	TRY variants differ in whether the petiole is included.
3115	Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole excluded	BERVO:0002369 Specific leaf area	TRY variants differ in whether the petiole, rhachis and midrib are included.
3116	Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole included	BERVO:0002369 Specific leaf area	TRY variants differ in whether the petiole, rhachis and midrib are included.
3117	Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): undefined if petiole is in- or exclu	BERVO:0002369 Specific leaf area	TRY variants differ in whether the petiole, rhachis and midrib are included.
3136	Root branching intensity 3: number of root tips per total root length	BERVO:0002542 Root tips per unit root length
3147	Root starch content per dry mass	BERVO:0002449 Root starch content per unit dry mass
3362	Photosynthesis: maximum rate of PEPc carboxylation (Vpmax) per leaf area	BERVO:0002593 Leaf maximum phosphoenolpyruvate carboxylation rate per unit area
3372	Seed nitrogen (N) content per seed	BERVO:0002616 Seed nitrogen content per seed
3373	Seed phosphorus (P) content per seed	BERVO:0002617 Seed phosphorus content per seed
3374	Leaf cell wall mass per leaf area	BERVO:0002605 Leaf cell wall mass per unit area
3375	Leaf cell wall mass per leaf dry mass	BERVO:0002606 Leaf cell wall content per unit dry mass
3379	Photosynthesis: intercellular CO2 concentration to ambient CO2 (ci/ca)	BERVO:0002561 Leaf intercellular to ambient carbon dioxide ratio
3381	Leaf surface area of mesophyll cells per leaf area	BERVO:0002600 Leaf mesophyll surface area exposed to intercellular space per unit leaf area
3382	Leaf surface area of chloroplasts exposed to intercellular airspace per leaf area	BERVO:0002601 Leaf chloroplast surface area exposed to intercellular space per unit leaf area
3383	Leaf mesophyll cell wall thickness	BERVO:0002603 Leaf mesophyll cell wall thickness
3393	Photosynthesis: quantum efficiency of photosystem II (maximum potential; FvFm)	BERVO:0002559 Leaf maximum quantum efficiency of photosystem II
3394	Photosynthesis quantum yield (QY; corresponding to photosynthetic efficiency)	BERVO:0002597 Leaf photosynthetic quantum yield
3400	Plant growth form simple consolidated	BERVO:0002378 Plant growth form	TRY variants differ in their value lists.
3401	Plant growth form detailed consolidated	BERVO:0002378 Plant growth form	TRY variants differ in their value lists.
3468	Leaf water & osmotic potential: leaf water potential at turgor loss point	BERVO:0002400 Leaf water potential at turgor loss point
3480	Root branching intensity: number of root branches per root length	BERVO:0001096 Root branching frequency
3482	Seed / ovule ratio	BERVO:0002578 Seed to ovule ratio
3483	Tree ring width	BERVO:0002443 Tree ring width
3485	Photosynthesis: rate of triose phosphate utilisation (TPU)	BERVO:0002594 Leaf triose phosphate utilisation rate per unit area
3515	Plant number of fruits per plant	BERVO:0002619 Fruit number per plant
3520	Litter decomposition: fraction of mass remaining	BERVO:0002493 Mass remaining fraction of decomposing plant material
3524	Litter decomposition: fraction of carbon (C) remaining	BERVO:0002494 Carbon remaining fraction of decomposing plant material
3525	Litter decomposition: fraction of nitrogen (N) remaining	BERVO:0002495 Nitrogen remaining fraction of decomposing plant material
3526	Litter decomposition: fraction of phosphorus (P) remaining	BERVO:0002496 Phosphorus remaining fraction of decomposing plant material
3553	Stem length per stem dry mass (stem specific length; SSL)	BERVO:0002442 Stem specific length
3603	Seed number per ground area	BERVO:0002572 Seed number per ground area
3613	Infructescence number of fruits (fruits per infructescence)	BERVO:0002581 Fruit number per infructescence
3614	Fruit number of seeds (seeds per fruit)	BERVO:0002430 Seed number per fruit
3634	Seed germination base temperature	BERVO:0002436 Seed germination base temperature
3635	Seed germination base water potential	BERVO:0002437 Seed germination base water potential
3636	Seed coat thickness	BERVO:0002574 Seed coat thickness
3660	Seed fresh mass	BERVO:0002426 Seed fresh mass
3719	Inflorescence number of flowers (floral density)	BERVO:0002580 Flower number per inflorescence
3753	Stem total bark thickness	BERVO:0002438 Bark thickness
3762	Stem total bark density	BERVO:0002439 Bark density
3764	Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: stem	BERVO:0002371 Stem specific density
3765	Stem relative total bark thickness	BERVO:0002529 Relative bark thickness
3767	Leaf equivalent water thickness (EWT)	BERVO:0002562 Leaf equivalent water thickness
3794	Stem hydraulic conductivity	BERVO:0002444 Stem hydraulic conductivity
3802	Photosynthesis: convexity parameter of light response curve	BERVO:0002598 Leaf photosynthetic light response curvature
3803	Photosynthesis: gross photosynthesis per leaf area	BERVO:0002595 Leaf gross photosynthetic rate per unit area
3804	Photosynthesis: gross photosynthesis per leaf dry mass	BERVO:0002596 Leaf gross photosynthetic rate per unit dry mass
3805	Photosynthesis: light saturation point	BERVO:0002555 Leaf light saturation point
3828	Leaf cuticle thickness	BERVO:0002550 Leaf cuticle thickness
3881	Inflorescence: number of fruits per inflorescence	BERVO:0002581 Fruit number per infructescence
3973	Photosynthesis: Quantum use efficiency	BERVO:0002597 Leaf photosynthetic quantum yield
3978	Photosynthesis: non-photochemical quenching (NPQ)	BERVO:0002599 Leaf non-photochemical quenching
3997	Wood compressive strength	BERVO:0002532 Wood compressive strength
3998	Wood flexural strength	BERVO:0002531 Wood flexural strength
3999	Wood tensile strength	BERVO:0002533 Wood tensile strength
4010	Stem specific hydraulic conductivity	BERVO:0002406 Sapwood-specific hydraulic conductivity
4024	Plot aboveground biomass per ground area	BERVO:0001971 Aboveground biomass
4025	Plot canopy height	BERVO:0001967 Canopy height
4029	Leaf vein length per leaf area	BERVO:0002480 Leaf vein density
4076	Fruit fresh mass	BERVO:0002435 Fruit fresh mass

Covered by a general term

BERVO term	TRY traits
BERVO:0001077 Root porosity	1786 Fine root porosity (volume of root air space); 2042 Fine root (absorptive) porosity (volume of root air space)
BERVO:0001970 Vegetation biomass	457 Coarse root dry mass per ground area; 798 Root dry mass per ground area; 1110 Seed or fruit biomass per ground area; 1501 Root necromass (dead dry mass) per ground area for specified soil depth increment; 1762 Coarse root necromass (dead dry mass) per ground area for specified soil depth increment; 1795 Fine root dry mass per ground area; 2018 Fine root necromass (dead dry mass) per ground area for specified soil depth increment; 2051 Fine root (absorptive) dry mass per ground area; 2307 Fine root (transport) dry mass per ground area; 2563 Belowground plant organ dry mass per ground area; 2786 Belowground plant organ necromass (dead dry mass) per ground area for specified soil depth increment
BERVO:0001971 Aboveground biomass	448 Branch dry mass per ground area; 494 Leaf dry mass per ground area; 524 Stem dry mass per ground area; 1094 Wood (sapwood) biomass per ground area; 1183 Stem heartwood biomass per ground area; 3124 Shoot: plant aboveground woody biomass per ground area
BERVO:0002055 Plant tissue carbon to nitrogen mass ratio	146 Leaf carbon/nitrogen (C/N) ratio; 150 Litter carbon/nitrogen (C/N) ratio; 165 Stem carbon/nitrogen (C/N) ratio; 166 Coarse root carbon/nitrogen (C/N) ) ratio; 167 Fine root carbon/nitrogen (C/N) ratio; 409 Shoot carbon/nitrogen (C/N) ratio; 432 Coarse woody debris (CWD): carbon/nitrogen (C/N) ratio; 694 Twig carbon/nitrogen (C/N) ratio; 830 Bark carbon/nitrogen (C/N) ratio; 842 Coarse woody debris (CWD) branch: carbon/nitrogen (C/N) ratio; 899 Litter (fine root) carbon/nitrogen (C/N) ratio; 945 Litter (leaf) carbon/nitrogen (C/N) ratio; 1055 Root carbon/nitrogen (C/N) ratio; 1059 Coarse woody debris (CWD) root: carbon/nitrogen (C/N) ratio; 1154 Coarse woody debris (CWD) stem: carbon/nitrogen (C/N) ratio; 1260 Wood carbon/nitrogen (C/N) ratio; 1545 Coarse root carbon/nitrogen (C/N) ratio; 2057 Fine root (absorptive) carbon/nitrogen (C/N) ratio; 2313 Fine root (transport) carbon/nitrogen (C/N) ratio; 2569 Belowground plant organ carbon/nitrogen (C/N) ratio; 2884 Litter (other than leaf: twigs, bark, flowers…) C/N ratio; 3703 Branch wood carbon/nitogen (C/N) ratio; 3711 Branch bark carbon/nitogen (C/N) ratio
BERVO:0002188 Leaf carbon content per unit dry mass	674 Leaf carbon (C) content per dry mass (cotyledons); 3955 Leaf carbon (C) content per dry mass (senescent leaves)
BERVO:0002189 Leaf nitrogen content per unit dry mass	667 Leaf nitrogen (N) content per dry mass (cotyledons); 3954 Leaf nitrogen (N) content per dry mass (senescent leaves)
BERVO:0002214 Delta carbon-13	89 Leaf carbon isotope signature (delta 13C); 90 Root carbon isotope signature (delta 13C); 829 Bark carbon (C) isotope signature (delta 13C); 1115 Shoot carbon (C) isotope signature (delta 13C); 1213 Wood carbon (C) isotope signature (delta 13C); 1252 Litter carbon (C) isotope signature (delta 13C); 3016 Wood carbon isotope signature (delta13C); 3707 Branch wood carbon isotope ratio (delta 13C); 3712 Branch bark carbon isotope ratio (delta 13C); 4048 Flower carbon isotope ratio (delta 13C); 4049 Stem carbon isotope ratio (delta 13C)
BERVO:0002216 Delta nitrogen-15	78 Leaf nitrogen (N) isotope signature (delta 15N); 81 Root nitrogen (N) isotope signature (delta 15N); 832 Bark nitrogen (N) isotope signature (delta 15N); 1001 Litter nitrogen (N) isotope signature (delta 15N); 1125 Shoot nitrogen (N) isotope signature (delta 15N); 1230 Wood nitrogen (N) isotope signature (delta 15N); 2036 Fine root (absorptive) nitrogen (N) isotope signature (delta 15N); 2292 Fine root (transport) nitrogen (N) isotope signature (delta 15N); 3708 Branch wood nitrogen isotope ratio (delta 15N); 3713 Branch bark nitrogen isotope ratio (delta 15N)
BERVO:0002217 Delta oxygen-18	3358 Leaf oxygen (O) isotope signature (delta 18O)
BERVO:0002223 Water delta oxygen-18	3540 Wood (sapwood) water oxygen isotope signature (delta18O)
BERVO:0002224 Water delta deuterium	3539 Wood (sapwood) water hydrogen isotope signature (delta2H)
BERVO:0002339 Plant tissue carbon content	72 Litter carbon (C) content per litter dry mass; 84 Root carbon (C) content per root dry mass; 407 Stem carbon (C) content per stem dry mass; 447 Branch carbon (C) content per branch dry mass; 456 Coarse root carbon (C) content per coarse root dry mass; 471 Fine root carbon (C) content per fine root dry mass; 617 Bark carbon (C) content per bark dry mass; 691 Twig carbon content per twig dry mass; 775 Shoot carbon (C) content per shoot dry mass; 841 Coarse woody debris (CWD) branch: carbon (C) content per branch CWD dry mass; 868 Coarse woody debris (CWD): carbon (C) content per CWD dry mass; 898 Litter (fine root) carbon (C) content per fine root litter dry mass; 944 Litter (leaf) carbon (C) content per leaf litter dry mass; 1058 Coarse woody debris (CWD) root: carbon (C) content per coarse root CWD dry mass; 1153 Coarse woody debris (CWD) stem: carbon (C) content per stem CWD; 1527 Coarse root carbon (C) content per coarse root dry mass; 2039 Fine root (absorptive) carbon (C) content per absorptive fine root dry mass; 2295 Fine root (transport) carbon (C) content per transport fine root dry mass; 2551 Belowground plant organ carbon (C) content per belowground plant organ dry mass; 2883 Litter (other than leaf: twigs, bark, flowers…) carbon (C) content per litter dry mass; 2944 Seed carbon (C) content per dry mass (or diaspore); 3368 Wood (sapwood) carbon (C) content per dry mass; 3369 Wood (heartwood) carbon (C) content per heartwood dry mass; 3709 Branch bark carbon (C) content per dry mass; 3790 Bark carbon (C) content per dry mass (alive bark); 3792 Bark: dead bark carbon (C) content per dry mass; 3923 Trap carbon (C) content per trap dry mass; 3957 Stem carbon (C) content per dry mass (senescent stem); 3959 Fruit (harvest organ) carbon (C) content per dry mass
BERVO:0002340 Plant tissue nitrogen content	70 Litter nitrogen (N) content per litter dry mass; 80 Root nitrogen (N) content per root dry mass; 339 Shoot nitrogen (N) content per shoot dry mass; 406 Stem nitrogen (N) content per stem dry mass; 450 Branch nitrogen (N) content per branch dry mass; 459 Coarse root nitrogen (N) content per coarse root dry mass; 475 Fine root nitrogen (N) content per fine root dry mass; 618 Bark nitrogen (N) content per bark dry mass; 684 Reproductive organ nitrogen (N) content per reproductive structures dry mass; 690 Twig nitrogen (N) content per twig dry mass; 854 Coarse woody debris (CWD) branch: nitrogen (N) content per branch CWD dry mass; 876 Coarse woody debris (CWD): nitrogen (N) content per CWD dry mass; 887 Litter (cone, strobilus) nitrogen (N) content per dry mass; 908 Litter (fine root) nitrogen (N) content per fine root litter dry mass; 954 Litter (leaf) nitrogen (N) content per leaf litter dry mass; 1165 Coarse woody debris (CWD) stem: nitrogen (N) content per stem CWD; 1199 Litter twig nitrogen (N) content per twig dry mass; 1229 Wood nitrogen (N) content per wood dry mass; 1262 Coarse woody debris (CWD) root: nitrogen (N) content per coarse root CWD dry mass; 2035 Fine root (absorptive) nitrogen (N) content per absorptive fine root dry mass; 2291 Fine root (transport) nitrogen (N) content per transport fine root dry mass; 2547 Belowground plant organ nitrogen (N) content per belowground plant organ dry mass; 2882 Litter (other than leaf: twigs, bark, flowers...) nitrogen (N) content per litter dry mass; 2945 Seed nitrogen (N) content per dry mass (or diaspore); 2987 Cone (strobilus) nitrogen (N) content per cone dry mass; 3376 Leaf cell wall nitrogen (N) content per cell wall dry mass; 3453 Wood (sapwood) nitrogen content per dry mass; 3455 Wood (heartwood) nitrogen (N) content per heartwood dry mass; 3704 Branch wood nitrogen (N) content per dry mass; 3710 Branch bark nitrogen (N) content per dry mass; 3791 Bark nitrogen (N) content per dry mass (alive bark); 3793 Bark: dead bark nitrogen (N) content per dry mass; 3922 Trap nitrogen (N) content per trap dry mass; 3956 Stem nitrogen (N) content per dry mass (senescent stem); 3958 Fruit (harvest organ) nitrogen (N) content per dry mass
BERVO:0002342 Belowground net primary production	1391 Root growth: root dry mass production per ground area per year; 1676 Coarse root growth: coarse root dry mass production per ground area per year; 1932 Fine root growth: fine root dry mass production per ground area per year; 2188 Fine root (absorptive) growth: absorptive fine root dry mass production per ground area per year; 2444 Fine root (transport) growth: transport fine root dry mass production per ground area per year
BERVO:0002344 Belowground litter production	1379 Root mortality: root dry mass mortality per ground area and time; 1664 Coarse root mortality: coarse root dry mass mortality per ground area and time; 1920 Fine root mortality: fine root dry mass mortality per ground area and time; 2176 Fine root (absorptive) mortality: absorptive fine root dry mass mortality per ground area and time
BERVO:0002352 Phenological event date	2956 Flower onset of flowering (first flowering date, flowering beginning)
BERVO:0002369 Specific leaf area	69 Litter SLA (leaf area per leaf dry mass, specific leaf area, 1/LMA of leaf litter)
BERVO:0002370 Leaf dry matter content	3055 Leaf dry mass per fresh mass (cotyledons, cotyledon LDMC)
BERVO:0002371 Stem specific density	1204 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: twigs and branches; 3452 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: heartwood; 3472 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: sapwood; 3473 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: branch; 3736 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: twig; 3877 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: branch
BERVO:0002374 Specific root length	614 Fine root length per fine root dry mass (specific fine root length, SRL); 1550 Coarse root length per coarse root dry mass (specific coarse root length, SRL); 2062 Fine root (absorptive) length per absorptive fine root dry mass (specific absorptive fine root lengt; 2318 Fine root (transport) length per transport fine root dry mass (specific transport fine root length,
BERVO:0002375 Root diameter	896 Fine root diameter; 1526 Coarse root diameter; 2038 Fine root (absorptive) diameter; 2294 Fine root (transport) diameter
BERVO:0002376 Root tissue density	1525 Coarse root tissue density (coarse root dry mass per coarse root volume); 1781 Fine root tissue density (fine root dry mass per fine root volume); 2037 Fine root (absorptive) tissue density (absorptive fine root dry mass per absorptive fine root volume; 2293 Fine root (transport) tissue density (transport fine root dry mass per transport fine root volume)
BERVO:0002380 Plant tissue phosphorus content	71 Litter phosphorus (P) content per litter dry mass; 340 Shoot phosphorus (P) content per shoot dry mass; 616 Stem phosphorus (P) content per stem dry mass; 619 Bark phosphorus (P) content per bark dry mass; 620 Coarse root phosphorus (P) content per coarse root dry mass; 683 Root phosphorus (P) content per root dry mass; 685 Reproductive organ phosphorus (P) content per reproductive structures dry mass; 698 Fine root phosphorus (P) content per fine root dry mass; 828 Twig phosporus (P) content per twig dry mass; 855 Coarse woody debris (CWD) branch: phosphorus (P) content per branch CWD dry mass; 863 Branch phosphorus (P) content per branch dry mass; 878 Coarse woody debris (CWD): phosphorus (P) content per CWD dry mass; 909 Litter (fine root) phosphorus (P) content per fine root litter dry mass; 957 Litter (leaf) phosphorus (P) content per leaf litter dry mass; 1068 Coarse woody debris (CWD) root: phosphorus (P) content per coarse root CWD dry mass; 1169 Coarse woody debris (CWD) stem: phosphorus (P) content per stem CWD; 2049 Fine root (absorptive) phosphorus (P) content per absorptive fine root dry mass; 2305 Fine root (transport) phosphorus (P) content per transport fine root dry mass; 2561 Belowground plant organ phosphorus (P) content per belowground plant organ dry mass; 2890 Litter other than leaf phosphorus (P) content per litter dry mass; 2988 Cone (strobilus) phosphorus (P) content per cone dry mass; 3000 Seed phosphorus (P) content per dry mass (or diaspore); 3924 Trap phosphorus (P) content per trap dry mass; 4011 Stem wood phosphorus (P) content per dry mass
BERVO:0002381 Plant tissue nitrogen to phosphorus mass ratio	56 Leaf nitrogen/phosphorus (N/P) ratio; 686 Stem nitrogen/phosphorus (N/P) ratio; 687 Root nitrogen/phosphorus (N/P) ratio; 688 Reproductive organ nitrogen/phosphorus (N/P) ratio; 3011 Fine root nitrogen/phosphorus (N/P) ratio; 3516 Litter nitrogen/phosphorus (N/P) ratio
BERVO:0002392 Leaf stomatal conductance	1265 Stomata conductance per leaf area: nighttime; 4032 Stomata conductance per leaf area: predawn; 4037 Stomata conductance per leaf area: midday
BERVO:0002396 Individual leaf dry mass	3050 Leaf dry mass (cotyledons)
BERVO:0002399 Leaf water potential	3541 Leaf water potential predawn; 3542 Leaf water potential midday
BERVO:0002411 Root dry matter content	695 Fine root dry mass per fine root fresh mass (fine root dry matter content; RDMC); 1538 Coarse root dry mass per coarse root fresh mass (coarse root dry matter content; RDMC)
BERVO:0002412 Root lignin content per unit dry mass	904 Litter (fine root) lignin content per fine root litter dry mass; 1064 Coarse woody debris (CWD) root: lignin content per coarse root CWD dry mass; 1528 Coarse root lignin content per coarse root dry mass; 1784 Fine root lignin content per fine root dry mass; 2040 Fine root (absorptive) lignin content per absorptive fine root dry mass
BERVO:0002413 Root respiration rate per unit dry mass	1535 Coarse root respiration rate per coarse root dry mass; 1791 Fine root respiration rate per fine root dry mass; 2047 Fine root (absorptive) respiration rate per absorptive fine root dry mass; 2303 Fine root (transport) respiration rate per transport fine root dry mass
BERVO:0002414 Root to shoot ratio	1778 Fine root/shoot ratio; 2034 Fine root (absorptive)/shoot ratio; 2546 Belowground plant organ/shoot ratio
BERVO:0002415 Root branching ratio	1841 Fine root branching intensity 1 (branching ratio): number of fine roots per higher order root; 2097 Fine root (absorptive) branching intensity 1 (branching ratio): number of absorptive fine roots per; 2353 Fine root (transport) branching intensity 1(branching ratio): number of transport fine roots per hig
BERVO:0002416 Specific root tip abundance	1843 Root tips per fine root dry mass (specific fine root tip abundance, SRTA); 2099 Root tips per absorptive fine root dry mass (specific absorptive fine root tip abundance, SRTA)
BERVO:0002417 Root lifespan	1809 Fine root persistence (lifespan, longevity); 2065 Fine root (absorptive) persistence (lifespan, longevity); 2321 Fine root (transport) persistence (lifespan, longevity)
BERVO:0002418 Plant rooting depth	1521 Coarse root rooting depth; 1777 Fine root rooting depth
BERVO:0002438 Bark thickness	3355 Bark thickness (inner bark); 3356 Bark thickness (outer bark); 3725 Twig total bark thickness; 3726 Twig inner bark thickness; 3754 Stem inner bark thickness; 3815 Twig outer bark thickness; 3844 Branch inner bark thickness; 3845 Branch outer bark thickness
BERVO:0002439 Bark density	3734 Twig total bark density; 3735 Twig inner bark density; 3763 Stem inner bark density; 3846 Stem outer bark density
BERVO:0002440 Stem dry matter content	3714 Wood dry mass per saturated wood fresh mass (wood dry matter content, WDMC)
BERVO:0002448 Root non-structural carbohydrate content per unit dry mass	1536 Coarse root total non-structural carbon (TNC) content per coarse root dry mass; 1792 Fine root total non-structural carbon (TNC) content per fine root dry mass; 2048 Fine root (absorptive) total non-structural carbon (TNC) content per absorptive fine root dry mass; 2304 Fine root (transport) total non-structural carbon (TNC) content per transport fine root dry mass
BERVO:0002449 Root starch content per unit dry mass	3187 Coarse root starch content per dry mass; 3224 Fine root starch content per dry mass; 3261 Fine root (absorptive) starch content per dry mass; 3298 Fine root (transport) starch content per dry mass
BERVO:0002450 Root stele diameter	1569 Coarse root stele diameter; 1825 Fine root stele diameter; 2081 Fine root (absorptive) stele diameter; 2337 Fine root (transport) stele diameter
BERVO:0002451 Root cortex thickness	1563 Coarse root cortex thickness; 1819 Fine root cortex thickness; 2075 Fine root (absorptive) cortex thickness; 2331 Fine root (transport) cortex thickness
BERVO:0002452 Root conduit diameter	1577 Coarse root conduit diameter; 1833 Fine root conduit diameter; 2089 Fine root (absorptive) conduit diameter; 2345 Fine root (transport) conduit diameter
BERVO:0002453 Specific root area	1557 Coarse root surface area per coarse root dry mass (specific coarse root area); 1813 Fine root surface area per fine root dry mass (specific fine root area); 2069 Fine root (absorptive) surface area per absorptive fine root dry mass (specific absorptive fine root; 2325 Fine root (transport) surface area per transport fine root dry mass (specific transport fine root ar
BERVO:0002454 Root turnover rate	1699 Coarse root dry mass turnover rate; 1955 Fine root dry mass turnover rate; 2211 Fine root (absorptive) dry mass turnover rate; 2467 Fine root (transport) dry mass turnover rate; 2723 Belowground plant organ dry mass turnover rate
BERVO:0002455 Plant tissue potassium content	44 Leaf potassium (K) content per leaf dry mass; 628 Stem potassium (K) content per stem dry mass; 646 Coarse root potassium (K) content per coarse root dry mass; 701 Shoot potassium (K) content per shoot dry mass; 856 Coarse woody debris (CWD) branch: potassium (K) content per branch CWD dry mass; 864 Branch potassium (K) content per branch dry mass; 879 Coarse woody debris (CWD): potassium (K) content per CWD dry mass; 910 Litter (fine root) potassium (K) content per fine root litter dry mass; 958 Litter (leaf) potassium (K) content per leaf litter dry mass; 1003 Litter potassium (K) content per litter dry mass; 1069 Coarse woody debris (CWD) root: potassium (K) content per coarse root CWD dry mass; 1086 Root potassium (K) content per root dry mass; 1170 Coarse woody debris (CWD) stem: potassium (K) content per stem CWD; 1810 Fine root potassium (K) content per fine root dry mass; 2066 Fine root (absorptive) potassium (K) content per absorptive fine root dry mass; 2322 Fine root (transport) potassium (K) content per transport fine root dry mass; 2578 Belowground plant organ potassium (K) content per belowground plant organ dry mass; 2891 Litter (other than leaf: twigs, bark, flowers...) potassium (K) content per litter dry mass; 2960 Bark potassium (K) content per bark dry mass; 2968 Twig potassium (K) content per dry mass; 2984 Cone (strobilus) potassium (K) content per cone dry mass; 3925 Trap potassium (K) content per trap dry mass; 4013 Stem wood potassium (K) content per dry mass
BERVO:0002456 Plant tissue calcium content	252 Leaf calcium (Ca) content per leaf dry mass; 623 Stem calcium (Ca) content per stem dry mass; 641 Coarse root calcium (Ca) content per coarse root dry mass; 704 Shoot calcium (Ca) content per shoot dry mass; 840 Coarse woody debris (CWD) branch: calcium (Ca) content per branch CWD dry mass; 867 Coarse woody debris (CWD): calcium (Ca) content per CWD dry mass; 897 Litter (fine root) calcium (Ca) content per fine root litter dry mass; 943 Litter (leaf) calcium (Ca) content per leaf litter dry mass; 991 Litter calcium (Ca) content per litter dry mass; 1054 Root calcium (Ca) content per root dry mass; 1057 Coarse woody debris (CWD) root: calcium (Ca) content per coarse root CWD dry mass; 1152 Coarse woody debris (CWD) stem: calcium (Ca) content per stem CWD; 1544 Coarse root calcium (Ca) content per coarse root dry mass; 1800 Fine root calcium (Ca) content per fine root dry mass; 2056 Fine root (absorptive) calcium (Ca) content per absorptive fine root dry mass; 2312 Fine root (transport) calcium (Ca) content per transport fine root dry mass; 2568 Belowground plant organ calcium (Ca) content per belowground plant organ dry mass; 2892 Litter (other than leaf: twigs, bark, flowers…) calcium (Ca) content per litter dry mass; 2957 Bark calcium (Ca) content per bark dry mass; 2965 Twig calcium (Ca) content per dry mass; 2973 Branch calcium (Ca) content per branch dry mass; 2981 Cone (strobilus) calcium (Ca) content per cone dry mass; 4012 Stem wood calcium (Ca) content per dry mass
BERVO:0002457 Plant tissue magnesium content	257 Leaf magnesium (Mg) content per leaf dry mass; 624 Stem magnesium (Mg) content per stem dry mass; 642 Coarse root magnesium (Mg) content per coarse root dry mass; 702 Shoot magnesium (Mg) content per shoot dry mass; 852 Coarse woody debris (CWD) branch: magnesium (Mg) content per branch CWD dry mass; 874 Coarse woody debris (CWD): magnesium (Mg) content per CWD dry mass; 906 Litter (fine root) magnesium (Mg) content per fine root litter dry mass; 952 Litter (leaf) magnesium (Mg) content per leaf litter dry mass; 997 Litter magnesium (Mg) content per litter dry mass; 1066 Coarse woody debris (CWD) root: magnesium (Mg) content per coarse root CWD dry mass; 1081 Root magnesium (Mg) content per root dry mass; 1162 Coarse woody debris (CWD) stem: magnesium (Mg) content per stem CWD; 1807 Fine root magnesium (Mg) content per fine root dry mass; 2063 Fine root (absorptive) magnesium (Mg) content per absorptive fine root dry mass; 2575 Belowground plant organ magnesium (Mg) content per belowground plant organ dry mass; 2893 Litter (other than leaf: twigs, bark, flowers...) magnesium (Mg) content per litter dry mass; 2961 Bark magnesium (Mg) content per bark dry mass; 2969 Twig magnesium (Mg) content per dry mass; 2976 Branch magnesium (Mg) content per branch dry mass; 2985 Cone (strobilus) magnesium (Mg) content per cone dry mass; 4014 Stem wood magnesium (Mg) content per dry mass
BERVO:0002458 Plant tissue sulfur content	262 Leaf sulfur (S) content per leaf dry mass; 629 Stem sulfur (S) content per stem dry mass; 647 Coarse root sulfur (S) content per coarse root dry mass; 859 Coarse woody debris (CWD) branch: sulfur (S) content per branch CWD dry mass; 912 Litter (fine root) sulfur (S) content per fine root litter dry mass; 961 Litter (leaf) sulfur (S) content per leaf litter dry mass; 1005 Litter sulfur (S) content per litter dry mass; 1088 Root sulfur (S) content per root dry mass; 1128 Shoot sulfur (S) content per shoot dry mass; 1174 Coarse woody debris (CWD) stem: sulfur (S) content per stem CWD; 1812 Fine root sulfur (S) content per fine root dry mass; 2068 Fine root (absorptive) sulfur (S) content per absorptive fine root dry mass; 2901 Litter (other than leaf: twigs, bark, flowers...) sulfur (S) content per litter dry mass; 2963 Bark sulfur (S) content per bark dry mass; 2971 Twig sulfur (S) content per dry mass; 2978 Branch sulfur (S) content per branch dry mass; 2989 Cone (strobilus) sulfur (S) content per cone dry mass
BERVO:0002459 Plant tissue sodium content	260 Leaf sodium (Na) content per leaf dry mass; 627 Stem sodium (Na) content per stem dry mass; 645 Coarse root sodium (Na) content per coarse root dry mass; 858 Coarse woody debris (CWD) branch: sodium (Na) content per branch CWD dry mass; 882 Coarse woody debris (CWD): sodium (Na) content per CWD dry mass; 911 Litter (fine root) sodium (Na) content per fine root litter dry mass; 960 Litter (leaf) sodium (Na) content per leaf litter dry mass; 1004 Litter sodium (Na) content per litter dry mass; 1071 Coarse woody debris (CWD) root: sodium (Na) content per coarse root CWD dry mass; 1087 Root sodium (Na) content per root dry mass; 1127 Shoot sodium (Na) content per shoot dry mass; 1173 Coarse woody debris (CWD) stem: sodium (Na) content per stem CWD; 1811 Fine root sodium (Na) content per fine root dry mass; 2067 Fine root (absorptive) sodium (Na) content per absorptive fine root dry mass; 2323 Fine root (transport) sodium (Na) content per transport fine root dry mass; 2898 Litter (other than leaf: twigs, bark, flowers...) sodium (Na) content per litter dry mass; 3464 Stem Na (sodium) content per dry mass
BERVO:0002460 Plant tissue manganese content	258 Leaf manganese (Mn) content per leaf dry mass; 703 Shoot manganese (Mn) content per shoot dry mass; 853 Coarse woody debris (CWD) branch: manganese (Mn) content per branch CWD dry mass; 875 Coarse woody debris (CWD): manganese (Mn) content per CWD dry mass; 907 Litter (fine root) manganese (Mn) content per fine root litter dry mass; 953 Litter (leaf) manganese (Mn) content per leaf litter dry mass; 1067 Coarse woody debris (CWD) root: manganese (Mn) content per coarse root CWD dry mass; 1163 Coarse woody debris (CWD) stem: manganese (Mn) content per stem CWD; 1269 Root manganese (Mn) content per root dry mass; 1559 Coarse root manganese (Mn) content per coarse root dry mass; 1815 Fine root manganese (Mn) content per fine root dry mass; 2071 Fine root (absorptive) manganese (Mn) content per absorptive fine root dry mass; 2327 Fine root (transport) manganese (Mn) content per transport fine root dry mass; 2897 Litter (other than leaf: twigs, bark, flowers...) manganese (Mn) content per litter dry mass; 2962 Bark manganese (Mn) content per bark dry mass; 2970 Twig manganese (Mn) content per dry mass; 2977 Branch manganese (Mn) content per branch dry mass; 2986 Cone (strobilus) manganese (Mn) content per cone dry mass; 2994 Stem manganese (Mn) content per dry mass; 3463 Stem manganese (Mn) (manganese) content per dry mass
BERVO:0002461 Plant tissue iron content	256 Leaf iron (Fe) content per leaf dry mass; 626 Stem iron (Fe) content per stem dry mass; 644 Coarse root iron (Fe) content per coarse root dry mass; 705 Shoot iron (Fe) content per shoot dry mass; 849 Coarse woody debris (CWD) branch: iron (Fe) content per branch CWD dry mass; 950 Litter (leaf) iron (Fe) content per leaf litter dry mass; 995 Litter iron (Fe) content per litter dry mass; 1077 Root iron (Fe) content per root dry mass; 1159 Coarse woody debris (CWD) stem: iron (Fe) content per stem CWD; 1804 Fine root iron (Fe) content per fine root dry mass; 2060 Fine root (absorptive) iron (Fe) content per absorptive fine root dry mass; 2572 Belowground plant organ iron (Fe) content per belowground plant organ dry mass; 2910 Litter (other than leaf: twigs, bark, flowers...) iron (Fe) content per litter dry mass; 2959 Bark iron (Fe) content per bark dry mass; 2967 Twig iron (Fe) content per dry mass; 2975 Branch iron (Fe) content per branch dry mass; 2983 Cone (strobilus) iron (Fe) content per cone dry mass
BERVO:0002462 Plant tissue zinc content	268 Leaf zinc (Zn) content per leaf dry mass; 638 Stem zinc (Zn) content per stem dry mass; 656 Coarse root zinc (Zn) content per coarse root dry mass; 860 Coarse woody debris (CWD) branch: zinc (Zn) content per branch CWD dry mass; 962 Litter (leaf) zinc (Zn) content per leaf litter dry mass; 1007 Litter zinc (Zn) content per litter dry mass; 1073 Coarse woody debris (CWD) root: zinc (Zn) content per coarse root CWD dry mass; 1092 Root zinc (Zn) content per root dry mass; 1129 Shoot zinc (Zn) content per shoot dry mass; 1259 Coarse woody debris (CWD) stem: zinc (Zn) content per stem CWD; 1814 Fine root zinc (Zn) content per fine root dry mass; 2070 Fine root (absorptive) zinc (Zn) content per absorptive fine root dry mass; 2326 Fine root (transport) zinc (Zn) content per transport fine root dry mass; 2964 Bark zinc (Zn) content per bark dry mass; 2972 Twig zinc (Zn) content per dry mass; 2979 Branch zinc (Zn) content per branch dry mass; 2990 Cone (strobilus) zinc (Zn) content per cone dry mass
BERVO:0002463 Plant tissue copper content	255 Leaf copper (Cu) content per leaf dry mass; 636 Stem copper (Cu) content per stem dry mass; 654 Coarse root copper (Cu) content per coarse root dry mass; 844 Coarse woody debris (CWD) branch: copper (Cu) content per branch CWD dry mass; 947 Litter (leaf) copper (Cu) content per leaf litter dry mass; 993 Litter copper (Cu) content per litter dry mass; 1074 Root copper (Cu) content per root dry mass; 1117 Shoot copper (Cu) content per shoot dry mass; 1803 Fine root copper (Cu) content per fine root dry mass; 2059 Fine root (absorptive) copper (Cu) content per absorptive fine root dry mass; 2315 Fine root (transport) copper (Cu) content per transport fine root dry mass; 2958 Bark copper (Cu) content per bark dry mass; 2966 Twig copper (Cu) content per dry mass; 2974 Branch copper (Cu) content per branch dry mass; 2982 Cone (strobilus) copper (Cu) content per cone dry mass
BERVO:0002464 Plant tissue aluminum content	249 Leaf aluminium (Al) content per leaf dry mass; 625 Stem aluminium (Al) content per stem dry mass; 643 Coarse root aluminium (Al) content per coarse root dry mass; 987 Litter aluminium (Al ) content per litter dry mass; 1049 Root aluminium (Al) content per root dry mass; 1113 Shoot aluminium (Al) content per shoot dry mass; 1149 Coarse woody debris (CWD) stem: aluminium (Al) content per stem CWD; 1797 Fine root aluminium (Al) content per fine root dry mass; 2053 Fine root (absorptive) aluminium (Al) content per absorptive fine root dry mass; 2913 Litter (other than leaf: twigs, bark, flowers…) aluminium (Al) content per litter dry mass
BERVO:0002465 Plant tissue nickel content	261 Leaf nickel (Ni) content per leaf dry mass; 635 Stem nickel (Ni) content per stem dry mass; 653 Coarse root nickel (Ni) content per coarse root dry mass; 1000 Litter nickel (Ni) content per litter dry mass; 1084 Root nickel (Ni) content per root dry mass; 1124 Shoot nickel (Ni) content per shoot dry mass; 1808 Fine root nickel (Ni) content per fine root dry mass; 2064 Fine root (absorptive) nickel (Ni) content per absorptive fine root dry mass; 2320 Fine root (transport) nickel (Ni) content per transport fine root dry mass
BERVO:0002466 Plant tissue silicon content	264 Leaf silicon (Si) content per leaf dry mass; 1270 Root silicon (Si) content per root dry mass; 1560 Coarse root silicon (Si) content per coarse root dry mass; 1816 Fine root silicon (Si) content per fine root dry mass; 3934 Stem silicon (Si) content per dry mass
BERVO:0002467 Plant tissue boron content	250 Leaf boron (B) content per leaf dry mass; 1318 Root boron (B) content per root dry mass; 1860 Fine root boron (B) content per fine root dry mass; 3555 Stem / wood boron (B) content per dry mass
BERVO:0002468 Plant tissue molybdenum content	259 Leaf molybdenum (Mo) content per leaf dry mass; 632 Stem molybdenum (Mo) content per stem dry mass; 650 Coarse root molybdenum (Mo) content per coarse root dry mass; 1083 Root molybdenum (Mo) content per root dry mass
BERVO:0002473 Individual leaf fresh mass	3053 Leaf fresh mass (cotyledons)
BERVO:0002476 Leaf lignin content per unit dry mass	951 Litter (leaf) lignin content per leaf litter dry mass
BERVO:0002477 Leaf cellulose content per unit dry mass	946 Litter (leaf) cellulose content per leaf litter dry mass
BERVO:0002483 Leaf transpiration rate per unit area	1267 Leaf transpiration rate at nighttime
BERVO:0002485 Plant tissue lignin content	73 Litter lignin content per litter dry mass; 425 Coarse woody debris (CWD): lignin content per CWD dry mass; 692 Twig lignin content per twig dry mass; 850 Coarse woody debris (CWD) branch: lignin content per branch CWD dry mass; 1122 Shoot lignin content per shoot dry mass; 1160 Coarse woody debris (CWD) stem: lignin content per stem CWD; 2552 Belowground plant organ lignin content per belowground plant organ dry mass; 2885 Litter (other than leaf: twigs, bark, flowers...) lignin content per litter dry mass; 4005 Wood lignin content per dry mass
BERVO:0002486 Plant tissue cellulose content	76 Litter cellulose content per litter dry mass; 91 Root cellulose content per root dry mass; 426 Coarse woody debris (CWD): cellulose content per CWD dry mass; 843 Coarse woody debris (CWD) branch: cellulose content per branch CWD dry mass; 900 Litter (fine root) cellulose content per fine root litter dry mass; 1060 Coarse woody debris (CWD) root: cellulose content per coarse root CWD dry mass; 1116 Shoot cellulose content per shoot dry mass; 1156 Coarse woody debris (CWD) stem: cellulose content per stem CWD; 1529 Coarse root cellulose content per coarse root dry mass; 1785 Fine root cellulose content per fine root dry mass; 2553 Belowground plant organ cellulose content per belowground plant organ dry mass; 2888 Litter (other than leaf: twigs, bark, flowers…) cellulose content per litter dry mass; 4006 Wood cellulose content per dry mass
BERVO:0002487 Plant tissue hemicellulose content	93 Root hemicellulose content per root dry mass; 94 Leaf hemi-cellulose content per leaf dry mass; 848 Coarse woody debris (CWD) branch: hemicellulose content per branch CWD dry mass; 903 Litter (fine root) hemicellulose content per fine root litter dry mass; 1121 Shoot hemicellulose content per shoot dry mass; 1312 Root hemicellulose content per root dry mass; 1598 Coarse root hemicellulose content per coarse root dry mass; 1854 Fine root hemicellulose content per fine root dry mass; 2622 Belowground plant organ hemicellulose content per belowground plant organ dry mass
BERVO:0002488 Plant tissue phenolics content	147 Leaf phenols content per leaf dry mass; 956 Litter (leaf) phenols content per leaf litter dry mass; 1002 Litter phenols content per litter dry mass; 1168 Coarse woody debris (CWD) stem: phenols content per stem CWD; 1271 Root phenols content per root dry mass; 1817 Fine root phenols content per fine root dry mass; 2073 Fine root (absorptive) phenols content per absorptive fine root dry mass; 2585 Belowground plant organ phenols content per belowground plant organ dry mass; 2896 Litter (other than leaf: twigs, bark, flowers...) phenols content per litter dry mass
BERVO:0002489 Plant tissue non-structural carbohydrate content	676 Stem total non-structural carbon (TNC) content per stem dry mass; 3732 Twig inner bark total non-structural carbohydrate concentration; 3733 Twig wood total non-structural carbohydrate concentration; 3746 Coarse root inner bark total non-structural carbohydrate concentration; 3747 Coarse root wood total non-structural carbohydrate concentration; 3760 Stem inner bark total non-structural carbohydrate concentration; 3761 Stem wood total non-structural carbohydrate concentration; 3975 Fibrous roots total nonstructural carbohydrate (TNC) content per dry mass
BERVO:0002490 Plant tissue starch content	3585 Seed starch content per seed dry mass; 3730 Twig inner bark starch concentration; 3731 Twig wood starch concentration; 3744 Coarse root inner bark starch concentration; 3745 Coarse root wood starch concentration; 3758 Stem inner bark starch concentration; 3759 Stem wood starch concentration
BERVO:0002491 Decomposition rate constant	846 Coarse woody debris (CWD) branch: decomposition rate constant; 902 Litter (fine root) decomposition rate constant; 949 Litter (leaf) decomposition rate constant; 1062 Coarse woody debris (CWD) root: decomposition rate constant; 1158 Coarse woody debris (CWD) stem: decomposition rate constant; 1519 Root debris decomposition rate constant; 1775 Coarse root debris decomposition rate constant; 1910 Litter (fine root) decomposition: decomposition rate constant; 2031 Fine root debris decomposition rate constant; 2287 Fine root (absorptive) debris decomposition rate constant; 2543 Fine root (transport) debris decomposition rate constant; 2799 Belowground plant organ debris decomposition rate constant; 2881 Litter (other than leaf: twigs, bark, flowers...) decomposition rate constant
BERVO:0002492 Plant tissue lignin to nitrogen mass ratio	851 Coarse woody debris (CWD) branch: lignin/nitrogen (lignin/N) ratio; 905 Litter (fine root) lignin/nitrogen (lignin/N) ratio; 1065 Coarse woody debris (CWD) root: lignin/nitrogen (lignin/N) ratio; 1161 Coarse woody debris (CWD) stem: lignin/nitrogen ratio; 1356 Root ratio of lignin to nitrogen (N); 1898 Fine root ratio of lignin to nitrogen (N); 2154 Fine root (absorptive) ratio of lignin to nitrogen (N); 2666 Belowground plant organ ratio of lignin to nitrogen (N); 2886 Litter (other than leaf: twigs, bark, flowers...) lignin/nitrogen (N) ratio
BERVO:0002493 Mass remaining fraction of decomposing plant material	1367 Root decomposition: fraction of root dry mass not decomposed to original root dry mass; 1653 Coarse root decomposition: fraction of coarse root dry mass not decomposed to original coarse root d; 1909 Litter (fine root) decomposition: fraction of fine root dry mass not decomposed to original fine roo; 2165 Fine root (absorptive) decomposition: fraction of absorptive fine root dry mass not decomposed to or; 2421 Fine root (transport) decomposition: fraction of transport fine root dry mass not decomposed to orig; 2677 Belowground plant organ decomposition: fraction of belowground plant organ dry mass not decomposed t
BERVO:0002494 Carbon remaining fraction of decomposing plant material	3148 Root decomposition: fraction of carbon (C) remaining; 3225 Fine root decomposition: fraction carbon (C) remaining; 3336 Belowground plant organ decomposition: fraction carbon (C) remaining
BERVO:0002495 Nitrogen remaining fraction of decomposing plant material	3149 Root decomposition: fraction of nitrogen (N) remaining; 3189 Coarse root decomposition: fraction nitrogen (N) remaining; 3226 Fine root decomposition: fraction nitrogen (N) remaining; 3263 Fine root (absorptive) decomposition: fraction nitrogen (N) remaining; 3300 Fine root (transport) decomposition: fraction nitrogen (N) remaining; 3337 Belowground plant organ decomposition: fraction nitrogen (N) remaining
BERVO:0002497 Plant tissue soluble sugar content	1891 Fine root water soluble sugar content per fine root dry mass; 2147 Fine root (absorptive) water soluble sugar content per absorptive fine root dry mass; 2403 Fine root (transport) water soluble sugar content per transport fine root dry mass; 2659 Belowground plant organ water soluble sugar content per belowground plant organ dry mass; 3728 Twig inner bark soluble sugar concentration; 3729 Twig wood soluble sugar concentration; 3742 Coarse root inner bark soluble sugar concentration; 3743 Coarse root wood soluble sugar concentration; 3756 Stem inner bark soluble sugar concentration; 3757 Stem wood soluble sugar concentration
BERVO:0002498 Plant tissue lignin carbon fraction	424 Fine root lignin carbon (C) content per fine root carbon (C) content; 1331 Root lignin carbon (C) content per root carbon (C) content; 2129 Fine root (absorptive) lignin carbon (C) content per absorptive fine root carbon (C) content; 2385 Fine root (transport) lignin carbon (C) content per transport fine root carbon (C) content
BERVO:0002499 Plant tissue carbon to phosphorus mass ratio	151 Leaf carbon/phosphorus (C/P) ratio; 3517 Root carbon/phosporus (C/P) ratio; 3518 Stem carbon/phosporus (C/P) ratio; 3519 Litter carbon/phosporus (C/P) ratio; 3899 Fine root carbon/phosporus (C/P) ratio
BERVO:0002500 Plant tissue nitrogen to potassium mass ratio	3500 Litter nitrogen/potassium (N/K) ratio; 3531 Leaf nitrogen/potassium (N/K) ratio; 3534 Root nitrogen/potassium (N/K) ratio; 3537 Stem nitrogen/potassium (N/K) ratio
BERVO:0002501 Plant tissue phosphorus to potassium mass ratio	3532 Leaf phosphorus/potassium (P/K) ratio; 3535 Root phosphorus/potassium (P/K) ratio; 3538 Stem phosphorus/potassium (P/K) ratio; 3544 Litter phosphorus/potassium (P/K) ratio
BERVO:0002502 Plant tissue carbon to potassium mass ratio	3530 Leaf carbon/potassium (C/K) ratio; 3533 Root carbon/potassium (C/K) ratio; 3536 Stem carbon/potassium (C/K) ratio; 3543 Litter carbon/potassium (C/K) ratio
BERVO:0002503 Plant tissue ash content	333 Leaf ash content per leaf dry mass; 758 Litter ash content per litter dry mass; 1324 Root ash content per root dry mass; 1866 Fine root ash content per fine root dry mass
BERVO:0002504 Plant tissue protein content	503 Leaf protein content per leaf dry mass; 959 Litter (leaf) protein content per leaf litter dry mass; 1171 Coarse woody debris (CWD) stem: protein content per stem CWD; 2908 Litter (other than leaf: twigs, bark, flowers...) protein content per litter dry mass; 3609 Shoot protein content per dry mass
BERVO:0002505 Plant tissue lipid content	1345 Root lipid content per root dry mass; 1887 Fine root lipid content per fine root dry mass; 2143 Fine root (absorptive) lipid content per absorptive fine root dry mass; 3979 Leaf lipid content per leaf dry mass
BERVO:0002506 Plant tissue tannin content	74 Litter tannin content per litter dry mass; 148 Leaf tannin content per leaf dry mass; 3015 Root tannin content per root dry mass
BERVO:0002507 Plant tissue construction cost	1856 Fine root construction cost per fine root dry mass; 2624 Belowground plant organ construction cost per belowground plant organ dry mass; 2953 Leaf construction cost per leaf dry mass
BERVO:0002508 Plant tissue acid-insoluble carbon fraction	1336 Root carbon (C) content not hydrolyzable in acid extractives per root carbon (C) content; 1622 Coarse root carbon (C) content not hydrolyzable in acid extractives per coarse root carbon (C) conte; 1878 Fine root carbon (C) content not hydrolyzable in acid extractives per fine root carbon (C) content; 2134 Fine root (absorptive) carbon (C) content not hydrolyzable in acid extractives per absorptive fine r; 2390 Fine root (transport) carbon (C) content not hydrolyzable in acid extractives per transport fine roo
BERVO:0002509 Plant tissue acid-insoluble residue content	1877 Fine root dry mass not hydrolyzable in acid extractives per fine root dry mass; 2133 Fine root (absorptive) dry mass not hydrolyzable in acid extractives per absorptive fine root dry ma
BERVO:0002510 Plant tissue pH	615 Leaf pH; 693 Twig pH; 699 Fine root pH; 1167 Coarse woody debris (CWD) stem: pH
BERVO:0002511 Root length production	1685 Coarse root growth: coarse root length production per ground area per year; 1941 Fine root growth: fine root length production per ground area per year; 2197 Fine root (absorptive) growth: absorptive fine root length production per ground area per year
BERVO:0002512 Root length mortality	1924 Fine root mortality: fine root length mortality per ground area and year; 2180 Fine root (absorptive) mortality: absorptive fine root length mortality per ground area and year; 2436 Fine root (transport) mortality: transport fine root length mortality per ground area and year
BERVO:0002513 Necromass to biomass ratio	1513 Root necromass to biomass ratio; 1774 Coarse root necromass to biomass ratio; 2030 Fine root necromass to biomass ratio; 2798 Belowground plant organ necromass to biomass ratio
BERVO:0002519 Plant tissue respiration rate per unit dry mass	442 Branch respiration rate per branch dry mass; 2559 Belowground plant organ respiration rate per belowground plant organ dry mass
BERVO:0002520 Vegetation nitrogen stock	451 Branch nitrogen (N) content per ground area; 455 Coarse root nitrogen (N) content per ground area; 468 Fine root nitrogen (N) content per ground area; 501 Crown (canopy) nitrogen (N) content per ground area; 528 Stem nitrogen (N) content per unit ground area; 1502 Root nitrogen (N) per ground area; 2787 Belowground plant organ nitrogen (N) per ground area
BERVO:0002521 Plant tissue water content	1097 Wood (sapwood) water content; 2887 Litter (other than leaf: twigs, bark, flowers...) water content per litter dry mass; 3120 Leaf water content per leaf dry mass (not saturated); 3554 Stem water cotent per stem dry mass
BERVO:0002522 Plant tissue saturated water content	3122 Leaf water content per leaf dry mass (at saturation); 3776 Flower saturated water content; 3780 Inflorescence saturated water content; 3863 Branch xylem water content per dry mass (saturated); 3864 Branch bark water content per dry mass (saturated)
BERVO:0002523 Wood vessel area fraction	3910 Stem wood cross-section: fraction of vessels
BERVO:0002524 Wood fibre area fraction	3683 Twig fibre fraction; 3913 Stem wood cross-section: fraction of fiber area (wall+lumen)
BERVO:0002525 Wood ray parenchyma area fraction	3670 Twig ray fraction; 3912 Stem wood cross-section: fraction of ray parenchyma; 4008 Stem wood cross-section: fraction of radial parenchyma
BERVO:0002526 Wood axial parenchyma area fraction	3669 Twig axial parenchyma fraction; 3911 Stem wood cross-section: fraction of axial parenchyma; 3915 Stem wood cross-section: fraction of axial parenchyma
BERVO:0002527 Wood parenchyma area fraction	3682 Twig axial+ray parenchyma fraction; 4007 Stem wood cross-section: fraction of total parenchyma (axial+radial)
BERVO:0002528 Plant tissue dry matter content	119 Plant biomass and allometry: Plant dry mass per plant fresh mass; 689 Twig dry mass per twig fresh mass (twig dry matter content, TDMC); 1010 Leaf petiole dry mass per petiole fresh mass (leaf petiole dry matter content; LDMC petiole); 3559 Flower dry matter content (flower dry mass per flower fresh mass); 3588 Shoot dry matter content per fresh mass; 3788 Inflorescence dry matter content (Inflorescence DMC)
BERVO:0002529 Relative bark thickness	3737 Twig relative total bark thickness; 3738 Twig relative inner bark thickness; 3766 Stem relative inner bark thickness; 3843 Stem relative outer bark thickness (outer bark thickness / radius)
BERVO:0002530 Plant tissue modulus of elasticity	3662 Twig modulus of elasticity (MOE); 3816 Twig modulus of elasticity (Youngs modulus); 3817 Twig wood modulus of elasticity (Youngs modulus); 3818 Twig bark modulus of elasticity (Youngs modulus)
BERVO:0002534 Root length per ground area	1768 Coarse root length density (RLD): coarse root length per ground area; 2024 Fine root length per ground area; 2280 Fine root (absorptive) length density (RLD): absorptive fine root length per ground area; 2536 Fine root (transport) length density (RLD): transport fine root length per ground area
BERVO:0002535 Root mass density	1757 Coarse root dry mass per soil volume; 2013 Fine root dry mass per soil volume
BERVO:0002536 Vegetation carbon stock	1498 Root carbon (C) content per ground area; 1761 Coarse root carbon (C) content per ground area; 2017 Fine root carbon content per ground area; 2273 Fine root (absorptive) carbon content per ground area; 2529 Fine root (transport) carbon (C) content per ground area
BERVO:0002537 Root area index	2012 Fine root area index (RAI): fine root surface area per ground area
BERVO:0002539 Root hair diameter	3229 Fine root root hair diameter; 3266 Fine root (absorptive) root hair diameter; 3303 Fine root (transport) root hair diameter
BERVO:0002540 Root stele diameter fraction	1568 Coarse root stele fraction: stele diameter divided by coarse root diameter; 1824 Fine root stele fraction: stele diameter divided by fine root diameter; 2080 Fine root (absorptive) stele fraction: stele diameter divided by absorptive fine root diameter; 2336 Fine root (transport) stele fraction: stele diameter divided by transport fine root diameter
BERVO:0002541 Root stele area fraction	1570 Coarse root cross-sectional fraction of stele area to coarse root area; 1826 Fine root cross-sectional fraction of stele area to fine root area; 2082 Fine root (absorptive) cross-sectional fraction of stele area to absorptive fine root area; 2338 Fine root (transport) cross-sectional fraction of stele area to transport fine root area
BERVO:0002542 Root tips per unit root length	3176 Coarse root branching intensity 3: number of root tips per total coarse root length; 3213 Fine root branching intensity 3: number of root tips per total fine root length; 3250 Fine root (absorptive) branching intensity 3: number of root tips per total fine root (absorptive) l
BERVO:0002543 Root acid phosphatase activity	1975 Fine root acid phosphatase activity per fine root dry mass; 2231 Fine root (absorptive) acid phosphatase activity per absorptive fine root dry mass
BERVO:0002544 Root length per soil volume	1769 Coarse root length per soil volume; 2025 Fine root length per soil volume; 2281 Fine root (absorptive) length per soil volume; 2537 Fine root (transport) length per soil volume
BERVO:0002561 Leaf intercellular to ambient carbon dioxide ratio	4034 Leaf ratio of intercellular to ambient CO2 concentration (CiCa) predawn; 4039 Leaf ratio of intercellular to ambient CO2 concentration (CiCa) midday
BERVO:0002563 Plant tissue bulk modulus of elasticity	3777 Flower bulk modulus of elasticity; 3784 Inflorescence bulk modulus of elasticity
BERVO:0002564 Plant tissue relative water content at turgor loss point	3783 Inflorescence relative water content at the point of turgor loss
BERVO:0002565 Plant tissue water content change to turgor loss point	3770 Leaf water content difference between full turgor and turgor loss, per dry mass; 3778 Flower water content difference between full turgor and turgor loss, per dry mass; 3786 Inflorescence water content difference between full turgor and turgor loss, per dry mass
BERVO:0002566 Plant tissue respiration temperature sensitivity	272 Leaf respiration rate in the dark temperature dependence; 445 Branch respiration temperature dependence; 516 Root respiration temperature dependence; 522 Stem respiration temperature dependence
BERVO:0002582 Plant organ mass fraction	110 Plant biomass and allometry: Leaf dry mass per plant dry mass (leaf weight ratio, LWR); 136 Plant biomass and allometry: Stem dry mass per plant dry mass; 137 Plant biomass and allometry: Branch dry mass per plant dry mass; 465 Plant biomass and allometry: Coarse root dry mass per plant dry mass; 466 Plant biomass and allometry: Fine root dry mass per plant dry mass; 470 Plant biomass and allometry: Root mass fraction (fraction of root dry mass per whole plant dry mass); 478 Plant biomass and allometry: Wood dry mass per plant dry mass; 1046 Plant biomass and allometry: Reproductive dry mass per plant dry mass (reproductive weight fraction); 1048 Plant biomass and allometry: Rhizome dry mass per plant dry mass (rhizome weight fraction); 1534 Plant biomass and allometry: Coarse root mass fraction (fraction of coarse root dry mass per whole p; 2005 Plant biomass and allometry: Fine root mass fraction (fraction of fine root dry mass to whole plant; 2261 Plant biomass and allometry: Fine root (absorptive) mass fraction (fraction of absorptive fine root; 2773 Plant biomass and allometry: Belowground plant organ mass fraction (fraction of belowground plant or; 3933 Plant biomass and allometry: root dry mass per plant dry mass (root mass ratio, root mass fraction); 4078 Leaf paracotyledon dry mass per plant dry mass; paracotyledon weight fraction
BERVO:0002583 Plant organ nitrogen fraction	669 Plant biomass and allometry: Root nitrogen (N) content per plant nitrogen (N) content; 670 Plant biomass and allometry: Stem nitrogen (N) content per plant nitrogen (N) content; 671 Plant biomass and allometry: Leaf nitrogen (N) content per plant nitrogen (N) content
BERVO:0002584 Root length ratio	2021 Plant biomass and allometry: Fine root length ratio (RLR): fine root length per plant dry mass
BERVO:0002592 Plant tissue respiration rate per unit nitrogen	500 Leaf respiration rate in the dark per leaf nitrogen (N) content
BERVO:0002593 Leaf maximum phosphoenolpyruvate carboxylation rate per unit area	4075 Photosynthesis C4: Vpmax25 per leaf area
BERVO:0002607 Plant tissue water potential	3774 Flower water potential; 3782 Inflorescence water potential
BERVO:0002608 Plant tissue osmotic potential	3773 Flower osmotic potential; 3781 Inflorescence osmotic potential
BERVO:0002609 Plant tissue hydraulic capacitance	711 Leaf water capacitance; 3775 Flower water capacitance; 3785 Inflorescence water capacitance
BERVO:0002610 Plant tissue osmotic solute content	3771 Leaf osmotically active compounds per dry mass; 3779 Flower osmotically active compounds per dry mass; 3787 Inflorescence osmotically active compounds per dry mass
BERVO:0002612 Plant relative growth rate	3459 Plant belowground relative growth rate

Related terms

TRY ID	TRY trait	BERVO term	Note
3	Leaf angle (inclination, orientation)	BERVO:0001632 Cosine of leaf angle	BERVO has only the cosine and sine of leaf angle, as canopy radiation parameters.
22	Photosynthesis pathway	BERVO:0000835 Plant photosynthetic type (C3 or C4)	The BERVO term distinguishes only C3 and C4; TRY includes CAM and intermediates.
39	Litter decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
49	Photosynthesis: intercellular CO2 concentration	BERVO:0000529 Leaf gaseous carbon dioxide concentration	EcoSIM's value for a plant functional type, as a concentration per unit volume rather than a mole fraction.
67	Stomata index (number of stomata per epidermis cells)	BERVO:0002469 Stomatal density	Stomata per epidermal cell, not per unit area.
125	Leaf area per leaf fresh mass (specific leaf area (SLA or 1/LMA) based on leaf fresh mass)	BERVO:0002369 Specific leaf area	Per unit fresh mass rather than dry mass.
138	Seed number per reproducton unit	BERVO:0002430 Seed number per fruit	Per reproduction unit, which TRY does not define; it may be a fruit, a flower or an inflorescence.
172	Stem conduit lumen cross-sectional area (vessels and tracheids) per stem sapwood cross-sectional are	BERVO:0002523 Wood vessel area fraction	Conduits, vessels with tracheids, as a lumen area per unit of sapwood area.
197	Plant functional type (PFT)	BERVO:8000785 Plant functional type	BERVO has Plant functional type as a concept, not a variable whose value is a PFT.
271	Photosynthesis light use efficiency (LUE)	BERVO:0002597 Leaf photosynthetic quantum yield	Light use efficiency is often a canopy or stand value, carbon fixed or biomass made per unit of radiation, rather than the initial slope of a leaf light response.
335	Plant reproductive phenology timing (flowering time)	BERVO:0002352 Phenological event date	Usually the months of flowering rather than a date.
336	Seed number per ramet	BERVO:0002618 Seed number per plant	Per ramet of a clonal plant rather than per genetic individual.
351	Seed number per dispersal unit	BERVO:0002430 Seed number per fruit	Per diaspore rather than per fruit; the two differ where a fruit splits into several diaspores.
371	Root vessel diameter	BERVO:0002452 Root conduit diameter	Vessels only; conduits include tracheids.
416	Leaf chlorophyll a/b ratio	BERVO:0002474 Leaf chlorophyll content per unit area	The ratio of chlorophyll a to chlorophyll b.
437	Plant number of leaves	BERVO:0000773 Leaf number	The BERVO term comes from a per-branch EcoSIM variable.
479	Photosynthesis CO2 compensation point	BERVO:0000542 Carbon dioxide compensation point	EcoSIM resolves it by node, not by leaf or plant.
504	Leaf Rubisco content per leaf area	BERVO:0000562 Leaf rubisco content	The BERVO term is per unit leaf mass, not per leaf area.
506	Leaf sugar content per leaf dry mass	BERVO:0002497 Plant tissue soluble sugar content	TRY says only sugar, which may include sugars that are not soluble.
515	Root respiration rate per root nitrogen (N) content	BERVO:0002413 Root respiration rate per unit dry mass	Per unit root nitrogen rather than dry mass.
517	Root respiration rate per root surface area	BERVO:0002413 Root respiration rate per unit dry mass	Per unit root surface area rather than dry mass.
518	Woody area index (woody stem area per ground area)	BERVO:0001880 Canopy stem area index	TRY does not say whether the woody area is projected, as in the BERVO term, or the total surface area.
520	Stem respiration rate per stem surface area	BERVO:0002441 Stem respiration rate per unit dry mass	Per unit stem surface area rather than dry mass.
521	Stem respiration rate per stem nitrogen (N) content	BERVO:0002441 Stem respiration rate per unit dry mass	Per unit stem nitrogen rather than dry mass.
532	Stem respiration rate per stem volume	BERVO:0002441 Stem respiration rate per unit dry mass	Per unit stem volume rather than dry mass.
534	Photosynthesis temperature dependence	BERVO:0002589 Leaf photosynthesis temperature sensitivity	TRY does not say what form the value takes; it may be a temperature optimum rather than a sensitivity.
587	Plant growth rate	BERVO:0002612 Plant relative growth rate	TRY does not say the rate is relative; it may be an absolute rate of mass or height growth.
662	Root sugar content per root dry mass	BERVO:0002497 Plant tissue soluble sugar content	TRY says only sugar, which may include sugars that are not soluble.
663	Leaf respiration rate in light per dry mass	BERVO:0002556 Leaf respiration rate in light per unit area	Per unit leaf dry mass rather than area.
664	Leaf respiration rate in light per leaf respiration rate in the dark	BERVO:0002556 Leaf respiration rate in light per unit area	The ratio of respiration in light to respiration in the dark.
710	Leaf water content at turgor loss point	BERVO:0002564 Plant tissue relative water content at turgor loss point	TRY says water content, not relative water content.
732	Leaf chlorophyll content per leaf nitrogen (N) content	BERVO:0002475 Leaf chlorophyll content per unit dry mass	Per unit leaf nitrogen rather than dry mass.
741	Leaf veins free ending density	BERVO:0002480 Leaf vein density	The density of free vein endings, not vein length per area.
751	Leaf epidermis conductance	BERVO:0002408 Leaf cuticular conductance	Epidermal conductance, which may include leakage through incompletely closed stomata as well as through the cuticle.
781	Stomata distribution to ab- and adaxial leaf surfaces	BERVO:0002469 Stomatal density	How the stomata are divided between the two leaf surfaces.
839	Bark thickness per stem diameter	BERVO:0002529 Relative bark thickness	Per stem diameter rather than radius, so half the value.
845	Coarse woody debris (CWD) branch: decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
869	Coarse woody debris (CWD): decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
901	Litter (fine root) decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
939	Leaf isoprene emission rate per leaf dry mass	BERVO:0002514 Leaf isoprene emission rate	Per unit leaf dry mass rather than area.
940	Leaf length excluding petiole (leaf lamina length)	BERVO:0002470 Leaf length	The length of the lamina alone, without the petiole.
948	Litter (leaf) decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
967	Leaf volatile organic carbon (VOC): monoterpene emission rate per leaf dry mass	BERVO:0002515 Leaf monoterpene emission rate	Per unit leaf dry mass rather than area.
974	Photosynthesis rate per leaf area: transition to TPU limited photosynthesis	BERVO:0002594 Leaf triose phosphate utilisation rate per unit area	The photosynthetic rate at which TPU starts to limit assimilation, not the TPU rate.
981	Leaf transpiration rate per leaf dry mass	BERVO:0002483 Leaf transpiration rate per unit area	Per unit leaf dry mass rather than area.
1061	Coarse woody debris (CWD) root: decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
1103	Seed number per flower	BERVO:0002430 Seed number per fruit	Per flower rather than per fruit; flowers that set no fruit count toward it.
1106	Seed number per stem	BERVO:0002618 Seed number per plant	Per stem rather than per plant.
1157	Coarse woody debris (CWD) stem: decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
1237	Wood vessel density	BERVO:0002410 Stem conduit density	Vessels only; conduits include tracheids.
1238	Wood vessel diameter	BERVO:0002409 Stem conduit diameter	Vessels only; conduits include tracheids.
1264	Leaf petiole specific length (petiole length per petiole dry mass)	BERVO:0002472 Petiole length	Petiole length per unit petiole dry mass.
1296	Root branching intensity 2: number of root tips per higher order root length	BERVO:0002542 Root tips per unit root length	Tips per unit length of the higher-order root that bears them, not per total root length.
1390	Root growth: root carbon (C) production per ground area and time	BERVO:0002342 Belowground net primary production	Carbon production, which the term's g{C} unit covers, but TRY gives no time base, so it may be a rate for part of a season rather than per year.
1392	Root growth: root dry mass production per ground area per month	BERVO:0002342 Belowground net primary production	Per month rather than per year, usually a rate for part of the season.
1393	Root growth: root dry mass production per ground area per duration of exposition	BERVO:0002342 Belowground net primary production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
1394	Root growth: root dry mass production per ground area per day	BERVO:0002342 Belowground net primary production	Per day rather than per year, usually a rate for part of the season.
1400	Root growth: root length production per ground area per exposition period	BERVO:0002511 Root length production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
1435	Mycorrhizal colonization: fraction of root length with mycorrhizla hyphae, arbuscules, or vesicles	BERVO:0002419 Arbuscular mycorrhizal root colonization	Counts root length with hyphae, arbuscules or vesicles; hyphae alone need not be arbuscular mycorrhizal.
1675	Coarse root growth: coarse root carbon (C) production per ground area and time	BERVO:0002342 Belowground net primary production	Carbon production, which the term's g{C} unit covers, but TRY gives no time base, so it may be a rate for part of a season rather than per year.
1678	Coarse root growth: coarse root dry mass production per ground area per duration of exposition	BERVO:0002342 Belowground net primary production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
1679	Coarse root growth: coarse root dry mass production per ground area per day	BERVO:0002342 Belowground net primary production	Per day rather than per year, usually a rate for part of the season.
1908	Fine root monthly decomposition rate	BERVO:0002491 Decomposition rate constant	A monthly rate; TRY does not say whether it is a rate constant or a mass loss.
1931	Fine root growth: fine root carbon (C) production per ground area and time	BERVO:0002342 Belowground net primary production	Carbon production, which the term's g{C} unit covers, but TRY gives no time base, so it may be a rate for part of a season rather than per year.
1933	Fine root growth: fine root dry mass production per ground area per month	BERVO:0002342 Belowground net primary production	Per month rather than per year, usually a rate for part of the season.
1934	Fine root growth: fine root dry mass production per ground area per duration of exposition	BERVO:0002342 Belowground net primary production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
1935	Fine root growth: fine root dry mass production per ground area per day	BERVO:0002342 Belowground net primary production	Per day rather than per year, usually a rate for part of the season.
1940	Fine root growth: fine root length production per ground area per exposition period	BERVO:0002511 Root length production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
1978	Fine root phosphatase activity per fine root dry mass	BERVO:0002543 Root acid phosphatase activity	Phosphatase or phosphomonoesterase activity without the acid pH stated.
1979	Fine root phosphatase activity per fine root length	BERVO:0002543 Root acid phosphatase activity	Per unit fine root length rather than dry mass.
2044	Fine root (absorptive) vessel diameter	BERVO:0002452 Root conduit diameter	Vessels only; conduits include tracheids.
2098	Fine root (absorptive) branching intensity 2: number of root tips per higher order root length	BERVO:0002542 Root tips per unit root length	Tips per unit length of the higher-order root that bears them, not per total root length.
2196	Fine root (absorptive) growth: absorptive fine root length production per ground area per exposition	BERVO:0002511 Root length production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
2214	Fine root (absorptive) turnover rate during growing season	BERVO:0002454 Root turnover rate	Over a growing season rather than a year.
2300	Fine root (transport) vessel diameter	BERVO:0002452 Root conduit diameter	Vessels only; conduits include tracheids.
2446	Fine root (transport) growth: transport fine root dry mass production per ground area per duration o	BERVO:0002342 Belowground net primary production	Over the exposure period of an ingrowth core or minirhizotron, whose length goes with the value, rather than a year.
2545	Belowground plant organ rooting depth	BERVO:0002418 Plant rooting depth	Belowground plant organs include rhizomes and tubers as well as roots, so the depth is not only of roots.
2703	Belowground plant organ growth: belowground plant organ dry mass production per ground area per day	BERVO:0002342 Belowground net primary production	Per day rather than per year, usually a rate for part of the season.
2880	Litter (other than leaf: twigs, bark, flowers...) decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
3001	Leaf chlorophyll content in SPAD units relative	BERVO:0002474 Leaf chlorophyll content per unit area	A relative optical index of chlorophyll, not a mass.
3006	Leaf intercellular airspace area per leaf area	BERVO:0002602 Leaf porosity	An area fraction on a cross-section, rather than a volume fraction.
3065	Branch cross-sectional fraction of parenchyma	BERVO:0002527 Wood parenchyma area fraction	A fraction of the whole branch cross-section, which may include bark and pith, rather than of the wood.
3066	Branch cross-sectional fraction of conduits (vessels)	BERVO:0002523 Wood vessel area fraction	A fraction of the whole branch cross-section, which may include bark and pith, rather than of the wood.
3067	Branch cross-sectional fraction of fiber	BERVO:0002524 Wood fibre area fraction	A fraction of the whole branch cross-section, which may include bark and pith, rather than of the wood.
3090	Plant time until death	BERVO:0002613 Plant lifespan	The time until death of a plant in a study, often under a treatment, rather than its lifespan.
3109	Leaf area (in case of compound leaves: leaflet, petiole excluded)	BERVO:0002368 Individual leaf area	The area of one leaflet of a compound leaf, not of the whole leaf.
3111	Leaf area (in case of compound leaves: leaflet, petiole included)	BERVO:0002368 Individual leaf area	The area of one leaflet of a compound leaf, not of the whole leaf.
3113	Leaf area (in case of compound leaves: leaflet, undefined if petiole is in- or excluded)	BERVO:0002368 Individual leaf area	The area of one leaflet of a compound leaf, not of the whole leaf.
3123	Seed germination rate per day (maximum)	BERVO:0002569 Seed germinability	The maximum germination rate per day, not the final fraction germinated.
3128	Photosynthesis rate per stomatal conductance	BERVO:0002482 Photosynthetic water use efficiency	Intrinsic water use efficiency, divided by stomatal conductance rather than transpiration.
3214	Fine root forks per fine root length.	BERVO:0002542 Root tips per unit root length	Forks rather than tips per unit root length.
3215	Fine root calcium (Ca) content per fine root fresh mass	BERVO:0002456 Plant tissue calcium content	Per unit fine root fresh mass rather than dry mass.
3216	Fine root potassium (K) content per fine root fresh mass	BERVO:0002455 Plant tissue potassium content	Per unit fine root fresh mass rather than dry mass.
3217	Fine root magnesium (Mg) content per fine root fresh mass	BERVO:0002457 Plant tissue magnesium content	Per unit fine root fresh mass rather than dry mass.
3218	Fine root phosphorus (P) content per fine root fresh mass	BERVO:0002380 Plant tissue phosphorus content	Per unit fine root fresh mass rather than dry mass.
3219	Fine root sulfur (S) content per fine root fresh mass	BERVO:0002458 Plant tissue sulfur content	Per unit fine root fresh mass rather than dry mass.
3220	Fine root aluminium (Al) content per fine root fresh mass	BERVO:0002464 Plant tissue aluminum content	Per unit fine root fresh mass rather than dry mass.
3221	Fine root manganese (Mn) content per fine root fresh mass	BERVO:0002460 Plant tissue manganese content	Per unit fine root fresh mass rather than dry mass.
3222	Fine root condensed tannins	BERVO:0002506 Plant tissue tannin content	Condensed tannins only, and TRY does not say per what.
3259	Fine root (absorptive) condensed tannins	BERVO:0002506 Plant tissue tannin content	Condensed tannins only, and TRY does not say per what.
3359	Leaf construction cost per leaf area	BERVO:0002507 Plant tissue construction cost	Per unit leaf area rather than dry mass.
3363	Stomata density ratio from one side of leaf to other	BERVO:0002469 Stomatal density	The ratio of the densities on the two leaf surfaces, not a density.
3384	Leaf mesophyll porosity (air space versus volume or area)	BERVO:0002602 Leaf porosity	The mesophyll only, and TRY allows an area fraction on a cross-section as well as a volume fraction.
3385	Leaf mesophyll conductance for CO2 per unit chloroplast surface area	BERVO:0002560 Leaf mesophyll conductance	Per unit chloroplast surface area rather than leaf area.
3390	Branch vessel density	BERVO:0002410 Stem conduit density	Vessels only, and in a branch; conduits include tracheids.
3391	Branch vessel diameter	BERVO:0002409 Stem conduit diameter	Vessels only, and in a branch; conduits include tracheids.
3398	Crown (canopy) carotenoid content per ground area (carotenoid mass per leaf area * LAI)	BERVO:0002545 Leaf carotenoid content per unit area	Per unit ground area for a canopy, leaf content times leaf area index.
3399	Crown (canopy) anthocyanin content per ground area (anthocyanin mass per leaf area * LAI)	BERVO:0002547 Leaf anthocyanin content per unit area	Per unit ground area for a canopy, leaf content times leaf area index.
3402	Leaf water content total	BERVO:0000176 Leaf water content	The BERVO term is water per unit leaf carbon, an EcoSIM parameter.
3474	Leaf chlorophyll content per leaf dry mass (Chl a)	BERVO:0002475 Leaf chlorophyll content per unit dry mass	Chlorophyll a only.
3475	Leaf chlorophyll content per leaf dry mass (Chl b)	BERVO:0002475 Leaf chlorophyll content per unit dry mass	Chlorophyll b only.
3476	Leaf chlorophyll content per leaf area (Chl a)	BERVO:0002474 Leaf chlorophyll content per unit area	Chlorophyll a only.
3477	Leaf chlorophyll content per leaf area (Chl b)	BERVO:0002474 Leaf chlorophyll content per unit area	Chlorophyll b only.
3479	Xylem hydraulic vulnerability curve (P20, P50, P80)	BERVO:0002398 Xylem vulnerability to embolism	A vulnerability curve, not a single water potential.
3521	Litter decomposition: carbon (C) content per litter dry mass	BERVO:0002339 Plant tissue carbon content	From litter decomposition studies, where the value may be taken at a stated time after the start; check the TRY data release before mapping it to the static tissue term.
3522	Litter decomposition: nitrogen (N) content per litter dry mass	BERVO:0002340 Plant tissue nitrogen content	From litter decomposition studies, where the value may be taken at a stated time after the start; check the TRY data release before mapping it to the static tissue term.
3523	Litter decomposition: phosphorus (P) content per litter dry mass	BERVO:0002380 Plant tissue phosphorus content	From litter decomposition studies, where the value may be taken at a stated time after the start; check the TRY data release before mapping it to the static tissue term.
3527	Litter decomposition: carbon/nitrogen (C/N) ratio	BERVO:0002055 Plant tissue carbon to nitrogen mass ratio	From litter decomposition studies, where the value may be taken at a stated time after the start; check the TRY data release before mapping it to the static tissue term.
3528	Litter decomposition: carbon/phosphorus (C/P) ratio	BERVO:0002499 Plant tissue carbon to phosphorus mass ratio	From litter decomposition studies, where the value may be taken at a stated time after the start; check the TRY data release before mapping it to the static tissue term.
3529	Litter decomposition: nitrogen/phosphorus (N/P) ratio	BERVO:0002381 Plant tissue nitrogen to phosphorus mass ratio	From litter decomposition studies, where the value may be taken at a stated time after the start; check the TRY data release before mapping it to the static tissue term.
3562	Root phosphomonoesterase activity	BERVO:0002543 Root acid phosphatase activity	Phosphatase or phosphomonoesterase activity without the acid pH stated.
3618	Fruits per ground area	BERVO:0002572 Seed number per ground area	Fruits rather than seeds per unit ground area.
3653	Leaf surface of chloroplasts area per leaf area	BERVO:0002601 Leaf chloroplast surface area exposed to intercellular space per unit leaf area	The name does not say the surface is the part exposed to the intercellular space; it may be the whole chloroplast surface.
3656	Leaf rubisco content per dry mass	BERVO:0000562 Leaf rubisco content	An EcoSIM parameter; the mass basis is not stated as dry mass.
3661	Twig leaf area per sapwood area ratio	BERVO:0002407 Huber value	The reciprocal, leaf area per sapwood area, for a twig.
3665	Twig hydraulically-weighted vessel diameter (Sperry equation)	BERVO:0002409 Stem conduit diameter	Vessels only, hydraulically weighted, and in a twig; conduits include tracheids.
3666	Twig vessel density (vessels per crossescional area)	BERVO:0002410 Stem conduit density	Vessels only, and in a twig; conduits include tracheids.
3667	Twig vessel lumen fraction	BERVO:0002523 Wood vessel area fraction	Vessel lumens only.
3795	Stem vessel lumen area	BERVO:0002445 Stem conduit lumen area	Vessels only; conduits include tracheids.
3796	Stem sap flow rate	BERVO:0001951 Sap flow	TRY does not give the basis of the rate; the BERVO term is per plant, and a sap flux density would be per unit sapwood area.
3834	Seed viability: fraction of seeds with low viabilty	BERVO:0002570 Seed viability	The fraction of seeds of low viability or without a live embryo, roughly the complement.
3835	Seed viability: fraction of seeds without embryo or dead embryo	BERVO:0002570 Seed viability	The fraction of seeds of low viability or without a live embryo, roughly the complement.
3850	Leaf width: left of midrib	BERVO:0002471 Leaf width	The width on one side of the midrib only.
3851	Leaf width: right of midrib	BERVO:0002471 Leaf width	The width on one side of the midrib only.
3852	Leaf width asymmetry	BERVO:0002471 Leaf width	The asymmetry of the two half-widths.
3856	Photosynthesis: quantum yield of photosystem 2	BERVO:0002559 Leaf maximum quantum efficiency of photosystem II	The operating efficiency of photosystem II in the light, not the maximum.
3857	Leaf protein content per area: proteomic analysis	BERVO:0002504 Plant tissue protein content	Per unit leaf area rather than dry mass, from a proteomic analysis.
3873	Leaf petiole length per petiole diameter	BERVO:0002472 Petiole length	Petiole length divided by petiole diameter.
3904	Leaf interveinal distance	BERVO:0002480 Leaf vein density	The distance between veins, which falls as vein density rises.
3906	Leaf anthoctanins in ACI units	BERVO:0002547 Leaf anthocyanin content per unit area	An optical anthocyanin content index (ACI), not a mass.
3932	Leaf chlorophyll index	BERVO:0002474 Leaf chlorophyll content per unit area	A relative optical index of chlorophyll, not a mass.
3945	Plant lifespan: age trees reach in forested stands	BERVO:0002613 Plant lifespan	The age trees reach in forested stands, rather than the lifespan of the species.
3951	Snag decomposition rate	BERVO:0002491 Decomposition rate constant	TRY does not say whether this is a rate constant or a mass loss per unit time.
3960	Plant biomass and allometry: leaf dry mass per shoot dry mass; leaf weight ratio, LWR aboveground	BERVO:0002582 Plant organ mass fraction	A fraction of the shoot dry mass, not of the whole plant.
3961	Plant biomass and allometry: stem dry mass per shoot dry mass; Stem mass fraction aboveground	BERVO:0002582 Plant organ mass fraction	A fraction of the shoot dry mass, not of the whole plant.
3962	Plant biomass and allometry: fruit (harvest organ) dry mass per shoot dry mass; Fruit mass fraction	BERVO:0002582 Plant organ mass fraction	A fraction of the shoot dry mass, not of the whole plant.
4009	Stem wood cross-section: fraction of vessel lumen	BERVO:0002523 Wood vessel area fraction	Vessel lumens only.
4033	Leaf intercellular CO2 concentration (Ci) predawn	BERVO:0000529 Leaf gaseous carbon dioxide concentration	The predawn intercellular concentration; EcoSIM's value is for a plant functional type, as a concentration per unit volume.
4046	Stem diameter growth	BERVO:0002346 Stem diameter increment	The BERVO term is the increment at breast height over a stated period.
4050	Leaf prophyll leaf length	BERVO:0002470 Leaf length	The length of a prophyll, the first leaf of a shoot.
4083	Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA) of total leaf area	BERVO:0002369 Specific leaf area	Uses the total leaf area rather than the one-sided area.

The plant-functional-traits schema

The schema harmonizes TRY, AusTraits and fine-root trait data into one record per taxon. It is preliminary, so these rows are where the next slice starts, not the limit of it. Several of its TRY columns are aggregates of more than one TRY trait. Leaf area, for example, is spread over seven traits that differ in how a compound leaf and its petiole are counted, so each slot lists every trait it may draw on. The leaf_mass_per_area slot has no row: TRY has no trait of that name, since LMA is the reciprocal of SLA, and BERVO already has Leaf mass per area (BERVO:0002185).

Schema slot	TRY trait	Status	BERVO term
diaspore_mass	26 Seed dry mass	match	BERVO:0002372 Seed dry mass
diaspore_mass	233 Dispersal unit dry mass	match	BERVO:0002373 Diaspore dry mass
growth_form	42 Plant growth form	match	BERVO:0002378 Plant growth form
growth_form	3400 Plant growth form simple consolidated	match	BERVO:0002378 Plant growth form
growth_form	3401 Plant growth form detailed consolidated	match	BERVO:0002378 Plant growth form
leaf_area	3108 Leaf area (in case of compound leaves: leaf, petiole excluded)	match	BERVO:0002368 Individual leaf area
leaf_area	3109 Leaf area (in case of compound leaves: leaflet, petiole excluded)	related	BERVO:0002368 Individual leaf area
leaf_area	3110 Leaf area (in case of compound leaves: leaf, petiole included)	match	BERVO:0002368 Individual leaf area
leaf_area	3111 Leaf area (in case of compound leaves: leaflet, petiole included)	related	BERVO:0002368 Individual leaf area
leaf_area	3112 Leaf area (in case of compound leaves: leaf, undefined if petiole in- or excluded)	match	BERVO:0002368 Individual leaf area
leaf_area	3113 Leaf area (in case of compound leaves: leaflet, undefined if petiole is in- or excluded)	related	BERVO:0002368 Individual leaf area
leaf_area	3114 Leaf area (in case of compound leaves undefined if leaf or leaflet, undefined if petiole is in- or e	match	BERVO:0002368 Individual leaf area
leaf_dry_matter_content	47 Leaf dry mass per leaf fresh mass (leaf dry matter content, LDMC)	match	BERVO:0002370 Leaf dry matter content
leaf_nitrogen_content	14 Leaf nitrogen (N) content per leaf dry mass	match	BERVO:0002189 Leaf nitrogen content per unit dry mass
leaf_type	43 Leaf type	match	BERVO:0002379 Leaf type
plant_height	18 Plant height	match	BERVO:0002365 Plant height
plant_height	3106 Plant height vegetative	match	BERVO:0002366 Vegetative plant height
plant_height	3107 Plant height generative	match	BERVO:0002367 Generative plant height
root_diameter	83 Root diameter	match	BERVO:0002375 Root diameter
root_diameter	896 Fine root diameter	broader	BERVO:0002375 Root diameter
root_nitrogen_content	80 Root nitrogen (N) content per root dry mass	broader	BERVO:0002340 Plant tissue nitrogen content
root_nitrogen_content	475 Fine root nitrogen (N) content per fine root dry mass	broader	BERVO:0002340 Plant tissue nitrogen content
root_tissue_density	82 Root tissue density (root dry mass per root volume)	match	BERVO:0002376 Root tissue density
root_tissue_density	1781 Fine root tissue density (fine root dry mass per fine root volume)	broader	BERVO:0002376 Root tissue density
specific_leaf_area	3115 Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole excluded	match	BERVO:0002369 Specific leaf area
specific_leaf_area	3116 Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole included	match	BERVO:0002369 Specific leaf area
specific_leaf_area	3117 Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): undefined if petiole is in- or exclu	match	BERVO:0002369 Specific leaf area
specific_root_length	614 Fine root length per fine root dry mass (specific fine root length, SRL)	broader	BERVO:0002374 Specific root length
specific_root_length	1080 Root length per root dry mass (specific root length, SRL)	match	BERVO:0002374 Specific root length
stem_specific_density	4 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density	match	BERVO:0002371 Stem specific density
stem_specific_density	3764 Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density: stem	match	BERVO:0002371 Stem specific density
woodiness	38 Plant woodiness	match	BERVO:0002377 Plant woodiness

Species attributes

TRY ID	TRY trait
200	Species occurrence range: number of floristic zones
218	Species occurrence range: native vs invasive
230	Species phyto-sociological group
242	Species occurrence range: mode of introduction
243	Species occurrence range: neophyt status
244	Species occurrence range: endemism
574	Species US Federal T/E Status
575	Species US Wetland Indicator
593	Species habitat characterisation / Plant requirement: soil texture
600	Species habitat characterisation / Plant requirement: soil pH
602	Species habitat characterisation / Plant requirement: precipitation
604	Plant human usage types
761	Species habitat characterisation / Plant requirement: soil type
788	Species light requirement
822	Species occurrence range: ecological occurrence range
823	Species occurrence range: continentality
824	Species occurrence range: invasiveness
825	Species occurrence range: climate type
826	Species occurrence range: altitude of origin
827	Species occurrence range: origin zonal
935	Leaf herbivore species
1130	Species environmental indicator value according to Ellenberg: continentality
1131	Species environmental indicator value according to Ellenberg: light
1132	Species environmental indicator value according to Ellenberg: moisture
1133	Species environmental indicator value according to Ellenberg: nitrogen
1134	Species environmental indicator value according to Ellenberg: pH
1135	Species environmental indicator value according to Ellenberg: salt tolerance
1136	Species environmental indicator value according to Ellenberg: temperature
1138	Species nutrient requirements
1139	Species occurrence rang: typical abundance where naturally occurring
1140	Species occurrence range characteristics
1141	Species occurrence range: elevation range
1142	Species soil pH requirement
1144	Species soil moisture requirements
1145	Species status (nativity at growth location)
2816	Species range: Highland or lowland vicariant of a genus
2942	Species important species diagnostic traits
3063	Species invasion introduction year
3091	Species habitat characterization: soil hydrology
3092	Species habitat characterization: precipitation
3093	Species habitat characterization: potential evapotranspiration
3094	Species habitat characterization: aridity index
3095	Species habitat characterization: habitat
3096	Species habitat characterization: vegetation type
3410	Species habitat characterization: substrate requirement
3545	Species phylogenetic line
3720	Species light preference
3885	Species habitat characterization: climate preference: presence in Koeppen-Geiger Climate Zone
3963	Species phylogenetic age
3965	Species use
4028	Plot dominant species
4042	Species phylogeny: number of species per genera